Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

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Ephemeroptera (Mayflies) of Michigan

Key to the Ephemeroptera of Michigan

Introduction

Although well surveyed in certain locations (especially by J. W. and F. A. Leonard), much work remains to adequately inventory our Michigan mayfly fauna, especially in the Upper Peninsula. The publication by Randolph and McCafferty (1998) is a useful step forward in documenting the general distribution of mayfly species in the Great Lakes. This work, in addition to records from Leonard's collections housed at the University of Michigan Museum of Zoology, Insect Division, forms the basis for this list. There are Michigan records for at least 126 species in 42 genera, with perhaps 41 additional species that have documented records nearby.

Many other records exist as synonymies; these are included under the current valid name, based on information located at the Mayfly Central web site. Genera are denoted in green; species denoted in bold blue have been recorded in Michigan; species denoted in black bold are likely to occur in Michigan based on their known distribution with nearby state/provincial localities from which certified records have been published. You can use the Find function in your web browser to locate families, genera, and species. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated.

    ACANTHAMETROPIDAE Edmunds, in Edmunds, Allen & Peters, 1963

      Acanthametropus Tshernova, 1948
        Acanthametropus pecatonica (Burks, 1953) Metreturus - IL, WI

    AMELETIDAE McCafferty, 1991

      Ameletus Eaton, 1885a
        Ameletus cryptostimulus Carle, 1978 - ON (McCafferty et al. 2008)
        Ameletus lineatus Traver, 1932 - Randolph and McCafferty 1998: 27
        Ameletus ludens Needham, 1905 - IN, OH (Randolph and McCafferty 1998: 28)

    AMETROPODIDAE Bengtsson, 1913

      Ametropus Albarda, 1878
        Ametropus fragilis Albarda, 1878 - Steven and Hilsenhoff 1979: 226 (as Ametropus neavei)
          (syn.) Ametropus neavei McDunnough, 1928

    ARTHROPLEIDAE Balthasar, 1937

      Arthroplea Bengtsson, 1909
        Arthroplea bipunctata (McDunnough, 1924c) - Randolph and McCafferty 1998: 29; Bright coll.

    BAETIDAE Leach, 1815

      Acentrella Bengtsson, 1913
        Acentrella parvula (McDunnough, 1932) - Yanoviak and McCafferty 1996: 203; (as Baetis armillatus)
          (orig.) Pseudocloeon parvulumMcDunnough, 1932a
          (comb.) Plauditus parvulus (McDunnough, 1932a)
          (syn.) Baetis armillatus McCafferty & Waltz, 1990
          (syn.) Plauditus armillatus McCafferty & Waltz, 1990
        Acentrella turbida (McDunnough), 1924 Pseudocloeon - Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 30-31
          (orig.) Pseudocloeon turbidum McDunnough, 1924a
          (syn.) Acentrella carolina (Banks, 1924)
          (syn.) Cloeon carolina (Banks, 1924)
          (syn.) Pseudocloeon carolina (Banks, 1924)

      Acerpenna Waltz & McCafferty, 1987
        Acerpenna macdunnoughi (Ide, 1937) Baetis (orig.) - Randolph and McCafferty 1998: 31-32
        Acerpenna pygmaea (Hagen, 1861) Cloe (orig.) - Leonard and Leonard 1962: 85 (as B. pygmaeus); Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 31 (as A. harti) and 32-33 (as A. pygmaea)
          (comb.) Baetis pygmaeus (Hagen, 1861)
          (syn.) Acerpenna sp. 1McCafferty & Davis, 1992
          (syn.) Acerpenna harti (McDunnough, 1924b)
          (syn.) Baetis harti McDunnough, 1924b
          (syn.) Baetis spiethi Berner, 1940

      Anafroptilum Kluge, 2011
        Anafroptilum album (McDunnough, 1926b) Centroptilum (orig.) - Leonard 1948: 5; Leonard and Leonard 1962: 83-84 (also as C. convexum); Randolph and McCafferty 1998: 48-49 (also as C. convexum and C. walshi)
          (comb.) Procloeon walshi (McDunnough, 1929)
          (syn.) Centroptilum convexum Ide, 1930
          (syn.) Centroptilum walshi McDunnough, 1929
          (syn.) Procloeon convexum Ide, 1930
          (syn.) Procloeon sp. 2 McCafferty & Davis, 1992
        Anafroptilum victoriae (McDunnough, 1938) Centroptilum (orig.) - N. A. Wiersema, UMMZINS-01528; also record from Marquette Co., near Ives Lake (Jacobus & Wiersema 2014: 388-389)

      Apobaetis Day, 1955: 126
        Apobaetis etowah (Traver, 1937) Pseudocloeon (orig.) - McCafferty 2009: 355 (43.0105N,-85.9362W, inferred from Grand River at Deer Creek inlet)
          (syn.) Apobaetis indeprensus Day, 1955: 127

      Baetis Leach, 1815
        Baetis brunneicolor McDunnough, 1925a - Leonard and Leonard 1962: 88 (and as B. hiemalis 88-90); Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 34-36
          (syn.) Baetis anachris Burks, 1953
          (syn.) Baetis hiemalis Leonard, 1950
          (syn.) Baetis phyllis Burks, 1953
        Baetis bundyae Lehmkuhl, 1973 - Dewalt & South 2015: 144 (locations 47.93491N -89.17479W and 47.93854N -89.15144W)
          (syn.) Baetis macani bundyae Lehmkuhl, 1973 (stat.)
        Baetis flavistriga McDunnough, 1921 - Leonard and Leonard 1962: 86-87 (as B. levitans); Kovalak 1978: 10 (as B. levitans); Kohler 1983: 1363; Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 37-38
          (syn.) Baetis caurinus Edmunds & Allen, 1957
          (syn.) Baetis levitans McDunnough, 1925b
          (syn.) Baetis nanus McDunnough, 1923
          (syn.) Baetis ochris Burks, 1953
          (syn.) Baetis sinuosus Navas, 1924
        Baetis intercalaris McDunnough, 1921 - Leonard and Leonard 1962: 90-91; Kovalak 1978: 10; Kohler 1983: 1363; Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 38-40
          (syn.) Baetis lasallei Banks, 1924
        Baetis phoebus McDunnough, 1923 - Leonard 1948: 5 (as B. cingulatus); Lyman 1956: 574; Leonard and Leonard 1962: 91 (may be syn. of B. flavistriga - compare Zhou et al. 2010 and Kluge
          (syn.) Baetis cingulatus McDunnough, 1925b
          (syn.) Baetis pallidula McDunnough, 1924b
          (syn.) Baetis pallidulus McDunnough, 1924b
          (syn.) Baetis quebecensis Hubbard, 1974
        Baetis pluto McDunnough, 1925b - Randolph and McCafferty 1998: 40
        Baetis tricaudatus Dodds, 1923 - Leonard and Leonard 1962: 85-86 (as B. vagans); Kovalak 1978: 10 (as B. vagans); Kohler 1983: 1363; Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 43-44
          (syn.) Baetis incertans McDunnough, 1925b
          (syn.) Baetis intermedius Dodds, 1923
          (syn.) Baetis jesmondensis McDunnough, 1938
          (syn.) Baetis moffati Dodds, 1923
          (syn.spell.) Baetis moffatti Dodds, 1923
          (syn.) Baetis sp. No. 1 Traver, 1932
          (syn.) Baetis vagans McDunnough, 1925b

      Callibaetis Eaton, 1881
        Callibaetis ferrugineus ferrugineus (Walsh, 1862) Cloe (orig.) - Needham 1908: 262-263; Lyman 1956: 574; Leonard and Leonard 1962: 83; Koss 1970: 98; Randolph and McCafferty 1998: 45-46
          (orig.) Cloe ferruginea Walsh, 1862
          (comb.) Baetis ferrugineus (Walsh, 1862)
          (comb.) Callibaetis ferrugineus (Walsh, 1862)
        Callibaetis fluctuans (Walsh, 1862) Cloe (orig.) - Randolph and McCafferty 1998: 46-47
          (comb.) Baetis fluctuans Walsh, 1862
          (syn.) Callibaetis traverae Upholt, 1937
        Callibaetis pallidus Banks, 1900 - McCafferty 2009: 355
          (syn.) Callibaetis semicostatus Banks, 1914
        Callibaetis pretiosus Banks, 1914 - N. Br. AuSable River nr. Lovells, Crawford Co. 24 May 1936; Paw Paw, Van Buren Co. 12 July 1940 INHS
        Callibaetis skokianus Needham, 1903 - Koss 1970: 98; Randolph and McCafferty 1998: 47
          (syn.) Callibaetis brevicostatus Daggy, 1945: 388
          (syn.) Baetis vagans McDunnough, 1925

      Cloeon Leach, 1815
        Cloeon dipterum (Linnaeus, 1761) - Oceana Co. (43.4851N, -86.3603W), coll. K.Perez, S.McCubbin, J.Cossey, A.Borisenko, data from Webb et al. 2012, as Cloeon cognatum.
          (comb.) Chloeon dipterum (Linnaeus, 1761)
          (comb.) Cloe diptera (Linnaeus, 1761)
          (syn.) Cloeon cognatum Stephens, 1835
          (syn.) Cloeon inscriptum Bengtsson, 1914
          (syn.) Cloeon pallida Leach, 1815
          (comb.) Cloeopsis diptera (Linnaeus, 1761)
          (orig.) Ephemera diptera Linnaeus, 1761

      Diphetor Waltz & McCafferty, 1987
        Diphetor hageni (Eaton, 1885a) - IL, IN, OH, WI (Randolph and McCafferty 1998: 50), expect in southern Michigan
          (comb.) Baetis hageni Eaton, 1885a
          (syn.) Baetis devinctus Traver, 1935
          (syn.) Baetis herodes Burks, 1953
          (syn.) Baetis parvus Dodds, 1923
          (syn.) Diphetor devinctus (Traver, 1935)
          (hom.) Cloe unicolor Hagen, 1861 (C. unicolor Curtis, 1834)
          (hom.comb.) Baetis unicolor (Hagen, 1861)

      Heterocloeon McDunnough, 1925a
        Heterocloeon ampla (Traver, 1932) Acentrella (orig.) - IN, OH, WI (Randolph and McCafferty 1998: 29-30 (as A. ampla)
          (comb.) Baetis amplus (Traver, 1932)
        Heterocloeon curiosum (McDunnough, 1923) Baetis - WI (Randolph and McCafferty 1998: 51), expect in UP on border with Wisconsin
          (orig.) Baetis curiosus (McDunnough, 1923)
          (comb.) Centroptilum curiosum McDunnough, 1923
          (syn.) Rheobaetis traverae Mller-Liebenau, 1974

      Iswaeon McCafferty & Webb, 2005
        Iswaeon anoka (Daggy, 1945: 391 as Pseudocloeon) - Leonard and Leonard 1962: 91-92 (as P. anoka); Koss 1970: 98; Randolph and McCafferty 1998: 48 (as Centroptilum alamance); McCafferty 2009: 356
          (comb.) Heterocloeon anoka (Daggy, 1945: 391)
          (syn.) Cloeon alamance Traver, 1932
          (syn.) Centroptilum alamance (Traver, 1932)
          (syn.) Pseudocloeon edmundsi Jensen, 1969

      Labiobaetis Novikova & Kluge, 1987
        Labiobaetis dardanus (McDunnough, 1923) Baetis (orig.) - IL, IN, OH, WI (Randolph and McCafferty 1998: 52)
          (comb.) Pseudocloeon dardanum (McDunnough, 1923)
          (syn.) Baetis elachistus Burks, 1953
        Labiobaetis ephippiatum (Traver, 1935) Baetis (orig.) - St. Clair River on Canadian side (McCafferty et al. 2008)
          (comb.) Pseudocloeon ephippiatum (Traver, 1935)
          (comb.spell.) Acentrella ephippiatus (Traver, 1935)
        Labiobaetis frondalis (McDunnough, 1925a) Baetis (orig.) - IL, IN, OH, WI (records on border with Wisconsin and Indiana; Randolph and McCafferty 1998: 53)
          (comb.) Pseudocloeon frondale (McDunnough, 1925)
          (syn.) Baetis australis Traver, 1932
          (syn.) Baetis baeticatus Burks, 1953
        Labiobaetis propinquus (Walsh, 1863) Baetis (orig.) - Leonard and Leonard 1962: 85 (as B. spinosus); Randolph and McCafferty 1998: 54-55; Bright coll. (River Styx, 46.8564N, -87.8447W, Marquette Co.)
          (comb. spell.) Acentrella propinquus (Walsh, 1863)
          (comb.) Pseudocloeon propinquum (Walsh, 1863)
          (renam.) Cloe propinqua Walsh, 1863
          (syn.) Baetis spinosus McDunnough, 1925a
          (syn.) Cloe vicina Walsh, 1862 (hom. C. vicina Hagen, 1861)

      Neocloeon Traver, 1932: 365
        Neocloeon alamance Traver, 1932: 365 - Randolph and McCafferty 1998: 48
          (comb.) Centroptilum alamance (Traver, 1932)
          (comb.) Cloeon alamance (Traver, 1932)
        Neocloeon triangulifer (McDunnough, 1931a) Cloeon (orig.) - McCafferty 2009: 356
          (syn.) Centroptilum sp. 1McCafferty & Davis, 1992

      <Paracloeodes Day, 1955: 121
        Paracloeodes fleeki McCafferty & Lenat 2003: 33 - OH, widespread (McCaffery and Lenat 2003: 34)
        Paracloeodes minutus (Daggy, 1945: 394 as Pseudocloeon minutum) - IN, WI, OH (Randolph and McCafferty 1998: 55)
          (syn.) Paracloeodes abditus Day, 1955: 122 (McCafferty & Waltz 1990)

      Plauditus Lugo-Ortiz & McCafferty, 1998
        Plauditus cestus (Provonsha & McCafferty, 1982) Pseudocloeon (orig.) - IL, IN, OH, ON, WI, widespread NWT-GA
          (comb.) Barbaetis cestus Provonsha & McCafferty, 1982
        Plauditus cingulatus (McDunnough, 1931) Pseudocloeon (orig.) - Leonard and Leonard 1962: 87 (as Baetis cinctutus); Randolph and McCafferty 1998: 36 (as Baetis cinctutus)
          (syn.) Baetis cinctutus McCafferty & Waltz, 1990
          (syn.) Plauditius cinctutus McCafferty & Waltz, 1990
        Plauditus dubius (Walsh, 1862) - Cloe (orig.)- Randolph and McCafferty 1998: 36-37 (as Baetis dubius)
          (comb.) Baetis dubius Walsh, 1862
          (comb.) Cloeon dubium Walsh, 1862
          (comb.) Pseudocloeon dubium Walsh, 1862
          (syn.) Cloeon chlorops McDunnough, 1923
          (syn.) Pseudocloeon chlorops McDunnough, 1923
        Plauditus punctiventris (McDunnough, 1923) Cloeon (orig.) - Leonard 1948: 5; Yanoviak and McCafferty 1996: 203 (as B. punctiventris) Randolph and McCafferty 1998: 41-42
          (comb.) Baetis punctiventris McDunnough, 1923
          (comb.) Pseudocloeon punctiventris (McDunnough, 1923)
          (syn.) Baetis sp. 1McCafferty & Davis, 1992
          (syn.) Psuedocloeon anoka Daggy, 1945
          (syn.) Pseudocloeon edmundsi Jensen, 1969
          (syn.) Pseudocloeon myrsum Burks, 1953
        Plauditus virilis (McDunnough, 1923) Cloeon - IN, OH (Randolph and McCafferty 1998: 45)
          (comb.spell.) Pseudocloeon virile, virilis McDunnough, 1923
          (comb.spell.) Baetis virile, virilis McDunnough, 1923
          (syn.) Acentrella sp. 1 Lugo-Ortiz & McCafferty, 1995
          (syn.) Acentrella sp. 2 Lugo-Ortiz & McCafferty, 1995

      Procleoeon Bengtsson, 1915
        Procloeon bellum (McDunnough, 1924a) Centroptilum - Randolph and McCafferty 1998: 56
        Procloeon fragile (McDunnough, 1923) Centroptilum - Randolph and McCafferty 1998: 56
        Procloeon mendax (Walsh, 1862) Cloeon - Randolph and McCafferty 1998: 56
        Procloeon pennulatum (Eaton, 1870) Centroptilum - Mackinaw Co. (Merritt et al. 2002); however, see Randolph and McCafferty 1998: 56-57
          (comb.) Pseudocentroptilum pennulatum Eaton, 1870
          (syn.) Centroptilum infrequens McDunnough, 1924a
        Procloeon quaesitum (McDunnough, 1931a) Centroptilum - IN (Randolph and McCafferty 1998: 57)
        Procloeon rivulare (Traver, 1935) Centroptilum - OH (McCafferty et al. 2008: 58)
        Procloeon rubropictum (McDunnough, 1923) Cloeon - Lyman 1956: 574 (as Cloeon rubropictum); Randolph and McCafferty 1998: 58
        Procloeon rufostrigatum (McDunnough, 1924a) Centroptilum (orig.) - Leonard and Leonard 1962: 84 (as C. rufostrigatum); Randolph and McCafferty 1998: 58
          (syn.) Centroptilum bistrigatum Daggy, 1945: 389
        Procloeon simplex (McDunnough, 1925a) Cloeon (orig.) - Randolph and McCafferty 1998: 58
        Procloeon viridoculare (Berner, 1940) Centroptilum (orig.) - Waltz and Munro 1996, Randolph and McCafferty 1998: 59

      Pseudocentroptiloides Jacob, 1986
        Pseudocentroptiloides usa Waltz & McCafferty, 1989 - McCafferty 2009: 357

      Waynokiops Hill, Pfeiffer & Jacobus, 2010
        Waynokiops dentatogriphus Hill, Pfeiffer & Jacobus, 2010 - IN (Steuben Co., ca. 8 km from Michigan border; Hill et al. 2010: 61-68)

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    BAETISCIDAE Banks, 1900 (Suborder Carapacea - armored mayflies)
    Key to the Baetiscidae of Michigan

      Baetisca Walsh, 1863
        Baetisca lacustrisMcDunnough, 1932a - IL, IN, OH, WI (expect in sw Michigan and western UP; see Randolph and McCafferty 1998: 61)
          (syn.) Baetisca bajkovi Neave, 1934
        Baetisca laurentinaMcDunnough, 1932a - Leonard 1948: 5; Leonard and Leonard 1962: 66; Randolph and McCafferty 1998: 62; Bright coll.
        Baetisca obesa (Say, 1839) Baetis (orig.) - Lyman 1956: 574; Leonard and Leonard 1962: 66; Koss 1970: 101; Randolph and McCafferty 1998: 62

    BEHNINGIIDAE Motas & Bacesco, 1937

      Dolania Edmunds & Traver, 1959
        Dolania americana Edmunds & Traver, 1959 - WI (Randolph and McCafferty 1998: 63)
        This species has not yet been recorded from Michigan, with regional records restricted to western Wisconsin (Mississippi River area). This species may be incountered in large rivers in sandy substrates, perhaps in the w LP).

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    CAENIDAE Newman, 1853

      Caenis Stephens, 1835
        Caenis amica Hagen, 1861 - Lyman 1956: 574 (as C. simulans); Leonard and Leonard 1962: 40-41 (as C. simulans); Koss 1970: 101 (as C. simulans); Randolph and McCafferty 1998: 65-66
          (syn.) Caenis simulans McDunnough, 1931
        Caenis anceps Traver, 1935 - Koss 1970: 101; Randolph and McCafferty 1998: 66-67
        Caenis diminuta Walker, 1853 - Needham 1908: 263; Randolph and McCafferty 1998: 67
        Caenis hilaris (Say, 1839b) Ephemera (orig.) - Cook Collection, MSU: Koss 1970:101; Randolph and McCafferty 1998: 67
        Caenis latipennis Banks, 1907 - Leonard and Leonard 1962: 41 (as C. jocosa, C. forcipata); Koss 1970: 101 (as C. forcipata and C. jocosa); Randolph and McCafferty 1998: 68-69
          (syn.) Caenis delicata Traver, 1935
          (syn.) Caenis forcipata McDunnough, 1931
          (syn.) Caenis gigas Burks, 1953
          (syn.) Caenis jocosa McDunnough, 1931
        Caenis punctata McDunnough, 1931 - Randolph and McCafferty 1998: 69
        Caenis tardata McDunnough, 1931 - Cook Collection, MSU - Koss 1969: 101
        Caenis youngi Roemhild, 1984 - Randolph and McCafferty 1998: 69-70

      Cercobrachys Soldán, 1986
        [Note: A specimen of Cercobrachys also has recently been collected, but its species identity has not yet been confirmed (Todd White, MSU, pers. comm.). The systematics of the family has recently been revised by Sun and McCafferty (2008); Michigan species previously included in Brachycercus are now placed to a new genus Sparbarus].
        Cercobrachys etowah Soldán, 1986 - IN, WI
        Cercobrachys fox Sun & McCafferty, 2008 - IN, WI
        Cercobrachys lilliei Sun & McCafferty, 2008 - WI
        Cercobrachys winnebago Sun & McCafferty, 2008 - WI

      Sparbarus Sun & McCafferty, 2008
        Sparbarus lacustris (Needham, 1918) - Caenis (orig.) - Lyman 1956: 574 (as Brachycercus lacustris); Leonard and Leonard 1962: 38 (as Brachycercus lacustris); Randolph and McCafferty 1998: 64 (as B. lacustris)
          (comb.) Brachycercus lacustris (Needham, 1918)
          (comb.) Eurycaenis lacustris (Needham, 1918)
          (syn.) Brachycercus idei (Lestage, 1931)
          (syn.) Brachycercus pallidus (Ide, 1930)
          (syn.) Eurycaenis idei Lestage, 1931
          (syn.) Eurycaenis pallida Ide, 1930
        Sparbarus maculatus (Berner, 1946) Brachycercus (orig.) - OH (Randolph and McCafferty 1998: 64)
          (syn.) Brachycercus pini Soldan, 1986
        Sparbarus nasatus (Soldan, 1986) Brachycercus (orig.) - IN, WI (Randolph and McCafferty 1998: 64)
        Sparbarus prudens (McDunnough, 1931) Eurycaenis (orig.) - IN, WI (expect in UP; Randolph and McCafferty 1998: 65)
          (comb.) Brachycercus prudens (McDunnough, 1931)

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    EPHEMERELLIDAE Klaplek, 1909

      Attenella Edmunds, 1971
        Attenella attenuata (McDunnough, 1925b) Ephemerella (orig.) - Allen and Edmunds 1961: (as E. attenuata); Koss 1970: 99 (as E. (Attenuatella) attenuata)
          (syn.) Ephemerella hirsuta Berner, 1946
        Attenella margarita (Needham, 1927) Ephemerella (orig.) - Yanoviak and McCafferty 1996: 203

      Dannella Edmunds, 1959
        Dannella lita (Burks, 1949) Ephemerella (orig.) - IL, IN, OH, ON (Randolph and McCafferty 1998: 87; McCafferty et al. 2008), may occur in extreme southern Michigan
          (comb.) Timpanoga lita (Burks, 1949)
        Dannella simplex (McDunnough, 1925b) Ephemerella (orig.) - Leonard 1948: 5 (as E. simplex); Leonard and Leonard 1962: 61-62 (as E. simplex); Allen and Edmunds 1962b: 334-336 (as E. simplex); Koss 1970: 99 (as E. (Dannella) simplex); Yanoviak and McCafferty 1996: 203 (as T. simplex)
          (comb.) Dannella simplex (McDunnough, 1925b)
          (comb.) Timpanoga simplex (McDunnough, 1925b)

      Drunella Needham, 1905
        Drunella cornuta (Morgan, 1911) Ephemerella (orig.) - Leonard and Leonard 1962: 47 (as E. cornuta); Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 71
          (syn.) Ephemerella depressa Ide, 1930
        Drunella cornutella (McDunnough, 1931a) Ephemerella (orig.) - Leonard 1948: 5 (as E. cornutella); Allen and Edmunds 1962a: 156-157; Yanoviak and McCafferty 1996: 203
        Drunella lata (Morgan, 1911) Ephemerella (orig.) - Leonard 1948: 5 (as E. lata); Leonard and Leonard 1962: 44-45 (as E. lata); Allen and Edmunds 1962a: 154-155 (as E. lata); Kovalak 1979:
          (syn.) Drunella longicornis (Traver, 1932)
          (syn.) Ephemerella inflata McDunnough, 1926b
          (syn.) Ephemerella longicornis Traver, 1932
        Drunella walkeri (Eaton, 1884) Baetis (orig., hom. B. fuscata Stephens, 1835) Ephemerella (orig.) - Leonard 1948: 5 (as E. fuscata); Leonard and Leonard 1962: 45-46 (as E. walkeri); Allen and Edmunds 1962a: 162-163 (as E. walkeri)
          (comb.) Drunella wayah (Traver, 1932)
          (hom.comb.) Ephemerella fuscata (Walker, 1853)
          (syn.) Ephemerella wayah Traver, 1932
          (syn.) Ephemerella bispina Needham, 1905

      Ephemerella Walsh, 1862
        Ephemerella aurivillii (Bengtsson, 1908) Chitonophora (orig.) - Allen and Edmunds 1965: 275-276
          (syn.) Cincticostella ezoensis (Gose, 1980)
          (spell.) Ephemerella aurivilliusi (Bengtsson, 1908) (spell.)
          (syn.) Ephemerella aroni Eaton, 1908
          (syn.) Ephemerella aronii Eaton, 1908
          (syn.) Ephemerella concinnata Traver, 1934
          (syn.) Ephemerella ezoensis Gose, 1980
          (syn.) Ephemerella nN Imanishi, 1940
          (syn.) Ephemerella norda McDunnough, 1924d
          (syn.) Ephemerella taeniata Tshernova, 1952
        Ephemerella catawba Traver, 1932 - WI
        Ephemerella dorothea dorthea Needham, 1908 - Leonard 1948: 5; Leonard and Leonard 1962: 52; Allen and Edmunds 1965: 258-259 (as E. dorothea); Yanoviak and McCafferty 1996: 203
          (syn.) Ephemerella dorothea Needham, 1908
        Ephemerella excrucians Walsh, 1862 - Needham 1908: 263 (Sta. 6: 42.5611N,-83.3370W; Sta. 26: 42.5665N,-83.3361W; Sta. 28: 42.5662N,-83.3360W); Leonard and Leonard 1962: 52; Allen and Edmunds 1965: 256
          (syn.) Ephemerella argo Burks, 1949
          (syn.) Ephemerella crenula Allen & Edmunds, 1965
          (spell.) Ephemerella excurcians Walsh, 1862
          (syn.) Ephemerella inermis Eaton, 1884
          (syn.) Ephemerella lacustris Allen & Edmunds, 1965
          (syn.) Ephemerella ora Burks, 1949
          (syn.) Ephemerella rama Allen, 1968
          (syn.) Ephemerella rossi Allen & Edmunds, 1965
          (syn.) Ephemerella semiflava McDunnough, 1926a
        Ephemerella invaria (Walker, 1853) Baetis (orig.) - Leonard 1948: 5 (and as E. rotunda); Leonard and Leonard 1962: 54, 56 (and as E. rotunda: 56, 59); Allen and Edmunds 1965: 263-264 (and as E. rotunda: 261-262); Kovalak 1978: 10
          (syn.) Chitonophora vernalis (Banks), 1914
          (syn.) Ephemerella choctawhatchee Berner, 1946
          (syn.spell.) Ephemerella femenina Needham, 1924
          (syn.) Ephemerella feminina Needham, 1924
          (syn.) Ephemerella fratercula McDunnough, 1925c
          (syn.) Ephemerella inconstans Traver, 1932
          (syn.) Ephemerella rotunda Morgan, 1911
          (syn.) Ephemerella simila Allen & Edmunds, 1965
          (syn.) Ephemerella vernalis Banks, 1914
        Ephemerella needhami (McDunnough, 1925a) - Leonard 1948: 5; Leonard and Leonard 1962: 50; Allen and Edmunds 1965: 274-275; Koss 1970: 100
        Ephemerella subvaria McDunnough, 1931a - Leonard 1948: 5; Leonard and Leonard 1962: 54; Allen and Edmunds 1965: 260-261; Kovalak 1978: 10; Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 79

      Eurylophella Tiensuu, 1935
        Eurylophella aestiva (McDunnough, 1931b) Ephemerella (orig.) - Randolph and McCafferty 1998: 80
        Eurylophella bicolor (Clemens, 1913) Ephemerella (orig.) - Lyman 1956: 574 (as Ephemerella bicolor); Allen and Edmunds 1963b: 603-604; Koss 1970: 100; Funk and Sweeney 1994: 245-248; Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 80-81
        Eurylophella funeralis (McDunnough, 1925c) Ephemerella (orig.) - Allen and Edmunds 1963b: 612-613; Koss 1970: 100; Randolph and McCafferty 1998: 81-82
        Eurylophella lutulenta (Clemens, 1913) Ephemerella (orig.) - Lyman 1956: 574 (as Ephemerella lutulenta); Leonard and Leonard 1962: 59 (as Ephemerella lutulenta); Randolph and McCafferty 1998: 82-83
        Eurylophella macdunnoughi Funk, in Funk & Sweeney 1994 - Randolph and McCafferty 1998: 83
        Eurylophella minimelia (McDunnough, 1931) Ephemerella (orig.) - Allen and Edmunds 1963b: 606-607
        Eurylophella temporalis (McDunnough, 1924c) Ephemerella (orig.) - Lyman 1956: 574 (as Ephemerella temporalis); Leonard and Leonard 1962: 61 (as Ephemerella temporalis); Koss 1970: 100; Randolph and McCafferty 1998: 84
        Eurylophella verisimilis (McDunnough, 1930) Ephemerella (orig.) - Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 85
          (spell.comb.) Eurylophella versimilis (McDunnough, 1930)

      Serratella Edmunds, 1959
        Serratella serrata (Morgan, 1911) Ephemerella (orig.) - Randolph and McCafferty 1998: 87
          (comb.) Serratella carolina (Berner & Allen, 1961)
          (comb.) Serratella sordida (McDunnough, 1925b)
          (syn.) Ephemerella carolina Berner & Allen, 1961
          (syn.) Ephemerella sordida McDunnough, 1925b
          (syn.) Ephemerella spiculosa Berner & Allen, 1961
          (syn.) Serratella spiculosa (McDunnough, 1925) and (Berner & Allen, 1961)
        Serratella serratoides (McDunnough, 1931a) Ephemerella (orig.) - Yanoviak and McCafferty 1996: 203

      Teloganopsis Ulmer, 1939
        Teloganopsis deficiens (Morgan, 1911) Ephemerella (orig.) - Leonard 1948: 5 (as E. deficiens); Leonard and Leonard 1962: 47, 50 (as E. deficiens); Allen and Edmunds 1963a: 587, 589 (as E. deficiens); Koss 1970: 100 (as E. (Serratella) deficiens); Kovalak 1978: 10 (as E. deficiens); Yanoviak and McCafferty 1996: 203 (as S. deficiens)
          (comb.) Serratella deficiens McDunnough, 1925
          (syn.) Ephemerella atrescens McDunnough, 1925b

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    EPHEMERIDAE Latreille, 1810

      Ephemera Linnaeus, 1758
        Ephemera blanda Traver, 1932 - ON (McCafferty et al. 2008: 446, record from the Bruce Peninsula in Ontario suggests it may also occur in Michigan)
          (syn.) Ephemera triplex Traver, 1935
        Ephemera simulans Walker, 1853 - Lyman 1956: 574; Leonard and Leonard 1962: 28; Koss 1970: 100; Randolph and McCafferty 1998: 89-90; Yanoviak and McCafferty 1996: 203; widespread in state, in many collections
          (syn.) Ephemera decora Walker, 1853
          (syn.) Ephemera natata (Walker, 1853)
          (syn.) Palingenia natata Walker, 1853
        Ephemera varia Eaton, 1883 - Needham 1908: 262; Needham 1920: 284

      Hexagenia Walsh, 1863
        Hexagenia atrocaudata McDunnough, 1924a - Spieth 1941: 240-242; Leonard and Leonard 1962: 30; Koss 1970: 100
        Hexagenia bilineata (Say, 1824a) Baetis - Needham 1908: 262; Needham 1920: 284; Randolph and McCafferty 1998: 91-92
          (comb.) Palingenia bilineata (Say, 1824)
        Hexagenia limbata (Serville, 1829) Ephemera (orig.) - Needham 1908: 262 (as H. variabiliis mis.spell.); Leonard 1948: 5 (as H. occulta); Hunt 1953; Spieth 1941: 250-253; Lyman 1956: 574-575 (also as H. affiliata?); Leonard and Leonard 1962: 30-33; Koss 1970: 100 (as H. limbata spp. limbata, occulta, and viridescens; as H. munda); widespread throughout the state
          (comb.) Palingenia limbata (Serville), 1829
          (syn.) Hexagenia affiliata McDunnough, 1927a
          (syn.) Hexagenia californica Upholt, 1937
          (syn.) Hexagenia carolina Traver, 1931
          (syn.) Hexagenia elegans Traver, 1931
          (syn.) Hexagenia kanuga Traver, 1937
          (syn.) Hexagenia marilandica Traver, 1931
          (syn.) Hexagenia mingo Traver, 1931
          (syn.) Hexagenia munda Eaton, 1883
          (syn.) Hexagenia occulta (Walker, 1853)
          (syn.) Hexagenia pallens Traver, 1935
          (syn.) Hexagenia rosacea Traver, 1931
          (syn.) Hexagenia variabilis Eaton, 1883
          (syn.) Hexagenia venusta Eaton, 1883
          (syn.) Hexagenia viridescens (Walker, 1853)
          (syn.) Hexagenia weewa Traver, 1931
          (syn.) Palingenia occulta Walker, 1853
          (syn.) Palingenia viridescens Walker, 1853
        Hexagenia rigida McDunnough, 1924a - Leonard and Leonard 1962: 33; Speith 1941: 267-269; Koss 1970: 100; Randolph and McCafferty 1998: 94
          (syn.nud.) Hexagenia form falcata Needham, 1920

      Litobrancha McCafferty, 1971
        Litobrancha recurvata Morgan, 1913 Hexagenia (orig.) - Speith 1941: 266-267; Leonard and Leonard 1962: 34; Koss 1970: 100; Bright coll.
          (spell.) Hexagenia recurvata Morgan, 1913

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    HEPTAGENIIDAE Needham in Needham & Betten, 1901

      Epeorus Eaton, 1881
        Epeorus namatus (Burks, 1946) Iron (orig.) - IN, OH (Randolph and McCafferty 1998: 96), may occur in small clear streams of southernmost LP of Michigan
        Epeorus suffusus (McDunnough, 1925a) Iron (orig.) -(J. Leonard record from UP)
        Epeorus vitreus (Walker, 1853) Palingenia (orig.) - Leonard 1948: 5 (as I. humeralis); Leonard and Leonard 1962: 118; Wotton et al. 1993: 2034; Yanoviak and McCafferty 1996: 203; Bright coll.
          (comb.) Heptagenia vitrea (Walker, 1853)
          (comb.) Iron vitreus (Walker, 1853)
          (comb.) Rhithrogena vitrea (Walker, 1853)
          (syn.) Epeorus humeralis Morgan, 1911
          (syn.) Epeorus rubidus (Traver, 1933)
          (syn.) Iron humeralis (Morgan, 1911)
          (syn.) Iron rubidus Traver, 1933

      Heptagenia Walsh, 1863
        Heptagenia diabasia Burks, 1946
        Heptagenia elegantula (Eaton, 1885b) Rhithrogena (orig.) - Leonard 1948: 5 (as H. hebe); Koss 1970: 99 (as H. hebe); Holomuzki 1996 (as H. hebe, per Randolph and McCaffery 1998: 97)
          (syn.) Heptagenia coxalis Banks, 1914
          (syn.) Heptagenia diabasia Burks, 1946
          (syn.) Heptagenia querula McDunnough, 1924a
        Heptagenia flavescens (Walsh, 1862) Palingenia (orig.) - Randolph and McCaffery 1998: 98-99
        Heptagenia pulla (Clemens, 1913) Heptagenia (orig.) - Leonard and Leonard 1962: 114-116; Koss 1970: 99; Yanoviak and McCafferty 1996: 203
          (comb.) Ecdyonurus pullus (Clemens, 1913)
          (spell.) Heptagenia pullus Clemens, 1913
          (nomen. nud.) Ecdyurus grandis Clemens, 1913

      Leucrocuta Flowers, 1980
        Leucrocuta aphrodite (McDunnough, 1926b) Heptagenia (orig.) - IL, IN, OH (Randolph and McCaffery 1998: 100-101, expect in s. LP)
          (comb.) Ecdyonurus aphrodite McDunnough, 1926b
        Leucrocuta hebe (McDunnough, 1924a) Heptagenia (orig.) - Leonard and Leonard 1962: 116; Holomuzki 1996 (as H. hebe, per Randolph and McCaffery 1998: 101-102); Yanoviak and McCafferty 1996: 203
          (comb.) Ecdyonurus hebe McDunnough, 1924
        Leucrocuta maculipennis (Walsh, 1863) Heptagenia (orig.) - IL, IN, OH, WI (Randolph and McCaffery 1998: 103, may occur in southernmost LP)
          (comb.) Ecdyonurus maculipennis Walsh, 1863
          (comb.) Ecdyurus maculipennis Walsh, 1863
        Leucrocuta walshi (McDunnough, 1926b) Heptagenia (orig.) - OH (Randolph and McCafferty 1998: 103-104, one records from Sandusky Co., may possibly occur in southeastermost LP)
          (comb.) Ecdyonurus walshi (McDunnough, 1926b)

      Maccaffertium Bednarik, 1979
        Maccaffertium exiguum (Traver, 1935) Stenonema (orig.) - Leonard and Leonard 1962: 109 (as S. integrum, mis.id., per Bednarik and McCafferty 1979: 18); Holomuzki 1996: 521; Randolph and McCafferty 1998: 115-116
          (comb.) Heptagenia exigua (Traver, 1933)
          (syn.) Stenonema alabamae Traver, 1937
          (syn.) Stenonema quinquespinum Lewis, 1974
          (syn.) Stenonema integrum, Leonard and Leonard (nec McDunnough, 1924) 1962
        Maccaffertium luteum (Clemens, 1913) Heptagenia (orig.) - Leonard and Leonard 1962: 109; Randolph and McCafferty 1998: 119
          (comb.) Ecdyonurus luteus Clemens, 1913
          (comb.) Stenonema luteum (Clemens, 1913)
        Maccaffertium mediopunctatum (McDunnough, 1926b) Ecdyonurus (orig.) - Leonard and Leonard 1962: 111 (as S. nepotellum); Koss 1970: 99 (as S. nepotellum); Kohler 1983: 1363 (as S. mediopunctatum)
          (comb.) Heptagenia mediopunctata McDunnough, 1926b
          (comb.) Heptagenia mediopunctata mediopunctata (McDunnough, 1926b)
          (comb.) Stenonema mediopunctatum McDunnough, 1926b
          (syn.) Ecdyonurus nepotellus McDunnough, 1933
          (syn.) Stenonema nepotellum McDunnough, 1933
        Maccaffertium mexicanum integrum (McDunnough, 1924b) Heptagenia (orig.) - IL, IN, OH, WI (Randolph and McCafferty 1998: 121-122
          (comb.) Heptagenia mexicana integer McDunnough, 1924b
          (comb.) Stenonema mexicanum integrum (McDunnough, 1924b)
          (comb.) Stenonema integrum (McDunnough, 1924)
          (syn.) Stenonema bellum Traver, 1933
          (syn.) Stenonema metriotes Burks, 1953
          (syn.) Stenonema wabasha Daggy, 1945: 378
        Maccaffertium modestum (Banks, 1910) Epeorus (orig.) - Leonard 1948: 5 (as S. rubrum); Leonard and Leonard 1962: 111-112 (as S. rubrum); Lewis 1974: 34 (as S. rubrum); Koss 1970: 99 (as S. rubromaculatum and S. rubrum); Kohler 1983: 1363; Yanoviak and McCafferty 1996: 203 (as S. modestum)
          (comb.) Heptagenia modesta (Banks, 1910)
          (comb.)Iron modestus (Banks, 1910)
          (comb.) Stenonema modestum (Banks, 1910)
          (syn.) Ecdyonurus ruber McDunnough, 1926b
          (syn.) Ecdyonurus rubromaculatus (Clemens, 1913)
          (syn.) Heptagenia rubromaculata Clemens, 1913
          (syn.) Stenonema annexum Traver, 1933
          (syn.) Stenonema ruber (McDunnough, 1926b)
          (syn.) Stenonema rubromaculatum (Clemens, 1913)
          (syn.) Stenonema rubrum (McDunnough, 1926)
          (syn.) Stenonema varium Traver, 1933
        Maccaffertium pulchellum (Walsh, 1862) Palingenia (orig.) - Needham 1908: 262; Leonard and Leonard 1962: 112, 114 (as S. pulchellum); Wiley and Kohler 1980: 619; Yanoviak and McCafferty 1996: 203 (as S. pulchellum)
          (id.) Heptagenia luridipennnis (Burmeister, 1839)
          (comb.) Ecdyonurus pulchellus (Walsh, 1862)
          (comb.) Heptagenia pulchella (Walsh, 1862)
          (comb.) Stenonema pulchellum (Walsh, 1862)
        Maccaffertium terminatum terminatum (Walsh, 1862) Palingenia (orig.) - Koss 1970: 99 (as S. lepton); Bednarik and McCafferty 1979: 40-43; Yanoviak and McCafferty 1996: 203 (as S. terminatum)
          (orig.) Palingenia terminata Walsh, 1862
          (comb.) Ecdyonurus terminatus (Walsh, 1862)
          (comb.) Heptagenia terminata (Walsh, 1862)
          (comb.) Heptagenia terminata terminata (Walsh, 1862)
          (comb.) Stenonema terminatum (Walsh, 1862)
          (comb.) Stenonema terminatum terminatum (Walsh, 1862)
          (syn.) Ecdyonurus bipunctatus McDunnough, 1926b
          (syn.) Stenonema ares Burks, 1953
          (syn.) Stenonema bipunctatum (McDunnough), 1926b
          (syn.) Stenonema lepton Burks, 1946
        Maccaffertium vicarium (Walker, 1853) Baetis (orig.) - Leonard 1948: 5 (as S. rivulicolum); Leonard and Leonard 1962: 104-108 (as Stenonema vicarium and S. fuscum); Koss 1970: 99 (as S. fuscum); Lewis 1974: 24 (as S. fuscum); Kohler 1983: 1363; Yanoviak and McCafferty 1996: 203 (as S. vicarium); Randolph and McCafferty 1998: 125-127
          (comb.) Ecdyonurus vicarius (Walker, 1853)
          (comb.) Ecdyurus vicarius (Walker, 1853)
          (comb.) Heptagenia vicaria (Walker, 1853)
          (comb.) Stenonema vicarium (Walker, 1853)
          (syn.) Baetis tessellata Walker, 1853
          (syn.) Stenonema tessellata (Walker, 1853)
          (syn.spell.) Ecdyonurus fusca (Clemens, 1913)
          (syn.) Ecdyonurus fuscum (Clemens, 1913)
          (syn.spell.) Ecdyonurus fuscus (Clemens, 1913)
          (syn.) Heptagenia fusca Clemens, 1913
          (syn.) Stenonema fuscum (Clemens, 1913)
          (syn.) Ecdyonurus rivulicolus McDunnough, 1933
          (syn.) Stenonema rivulicolum (McDunnough, 1933)

      Nixe Flowers, 1980
        Nixe inconspicua (McDunnough, 1924a) Heptagenia (orig.) - IL, IN, OH (Randolph and McCafferty 1998: 105, may occur in s. LP)
          (comb.) Ecdyonurus inconspicuus McDunnough, 1924
        Nixe lucidipennis (Clemens, 1913) Ecdyurus (orig.) - Lyman 1956: 574
          (comb.) Ecdyonurus lucidipennis Clemens, 1913
          (comb.) Heptagenia lucidipennis Clemens, 1913
        Nixe perfida (McDunnough, 1926a) Heptagenia (orig.) - IL, IN, OH (Randolph and McCafferty 1998: 106-107)
          (comb.) Ecdyonurus perfidus McDunnough, 1926

      Rhithrogena Eaton, 1881
        Rhithrogena impersonata (McDunnough, 1925a) Heptagenia (orig.) - Leonard 1948: 5; Leonard and Leonard 1962: 120-121 (and as R. sanguinea); Yanoviak and McCafferty 1996: 203
          (syn.) Rhithrogena sanguinea Ide, 1954
        Rhithrogena jejuna Eaton, 1885b - Leonard and Leonard 1962: 119-120
          (hom.) Baetis fusca Walker, 1853 (B. fusca Burmeister, 1839)
          (hom.comb.) Heptagenia fusca (Walker, 1853)
          (hom.comb.) Rhithrogena fusca (Walker, 1853)
        Rhithrogena manifesta Eaton, 1885b - Leonard and Leonard 1962: 119 (as R. pellucida); Yanoviak and McCafferty 1996: 203 (as R. pellucida)
          (comb.) Heptagenia manifesta (Eaton, 1885b)
          (syn.) Rhithrogena pellucida Daggy, 1945: 383
        Rhithrogena undulata (Banks, 1924) Epeorus - Yanoviak and McCafferty 1996: 203

      Stenacron Jensen, 1974
        Stenacron candidum (Traver, 1935) Stenonema (orig.) - IL, IN, OH (Randolph and McCafferty 1998: 110; record in northern Indiana make this species possible in southernmost LP)
          (comb.) Heptagenia candida (Traver, 1935)
        Stenacron gildersleevei (Traver, 1935) Stenonema (orig.) - IL, IN, OH, ON (McCafferty et al. 2008; however, see Koss 1970: 99 - check record at MSU)
          (comb.) Heptagenia gildersleevei (Traver, 1935)
        Stenecron minnetonka (Daggy, 1945: 376 as Stenonema) - MN, IL, IN, OH (Randolph and McCafferty 1998: 114; one record in Lake Co., IN, may occur in sw LP; however, see Koss 1970: 99 - check record at MSU)
          (comb.) Heptagenia minnetonka (Daggy, 1945)
        Stenacron interpunctatum (Say, 1839) Baetis (orig.) - Lyman 1945; 1956: 574; Leonard and Leonard 1962: 99-103 (also as Stenonema canadense, S. frontale, and S. heterotarsale); Koss 1970: 99 (as S. interpunctatus, S. canadense, S. frontale and S. heterotarsale); Wiley and Kohler 1980: 619; Kohler 1983: 1363; Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 111-114; widespread in state, in many collections
          (comb.) Stenonema interpunctatum (Say, 1839)
          (comb.) Heptagenia interpunctata (Say, 1839)
          (comb.) Palingenia interpunctata (Say, 1839)
          (syn.) Baetis canadensis Walker, 1853
          (syn.) Ecdyonurus canadensis (Walker, 1853)
          (syn.) Ecdyurus canadensis (Walker, 1853)
          (syn.) Heptagenia canadensis (Walker, 1853)
          (syn.) Stenacron canadense (Walker, 1853)
          (syn.) Stenonema canadense (Walker, 1853)
          (syn.) Ecdyonurus frontalis (Banks, 1910)
          (syn.) Stenacron frontale (Banks, 1910)
          (syn.) Ecdyonurus heterotarsalis McDunnough, 1933
          (syn.) Heptagenia frontalis Banks, 1910
          (syn.) Stenonema frontale (Banks, 1910)
          (syn.) Stenacron areion (Burks, 1953)
          (syn.) Stenonema areion Burks, 1953
          (syn.) Stenacron heterotarsale (McDunnough, 1933)
          (syn.) Stenonema heterotarsale (McDunnough, 1933)
          (syn.) Stenonema affine Traver, 1933
          (syn.) Stenonema conjunctum Traver, 1935
          (syn.) Stenonema majus Traver, 1935
          (syn.) Stenonema ohioense Traver, 1935
          (syn.) Stenonema proximum Traver, 1935

      Stenonema Traver, 1933
        Stenonema femoratum (Say, 1823) Baetis (orig.) - Lyman 1945; 1956: 574; Leonard and Leonard 1962: 103-104 (as S. tripuncatatum); Koss 1970: 99 (as S. tripunctatum); Bednarik and McCafferty 1979: 13-17; Duffy et al. 1987: 49 (as S. tripunctatum); Randolph and McCafferty 1998: 117-118
          (comb.) Ecdyonurus femoratus (Say, 1823)
          (comb.) Heptagenia femorata (Say, 1823)
          (comb.) Siphlurus femoratus (Say, 1823)
          (syn.) Ecdyonurus tripunctatus (Banks, 1910) (syn)
          (syn.) Heptagenia tripunctata Banks, 1910
          (syn.) Stenonema birdi Traver, 1935
          (syn.) Stenonema scitulum Traver, 1935
          (syn.) Stenonema tripunctatum (Banks, 1910)

      Spinadis Edmunds & Jensen, 1974
        Spinadis simplex (Walsh, 1863) Heptagenis (orig.) - IN, WI
          (comb.) Anepeorus simplex (Walsh, 1863)
          (syn.) Spinadis wallacei Edmunds & Jensen, 1974

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    ISONYCHIIDAE Burks, 1953

      Isonychia Eaton, 1871
        Isonychia bicolor (Walker, 1853) Palingenia (orig.) - Leonard 1948: 5 (as I. albomanicata); Leonard and Leonard 1962: 80, 82 (and as I. sadleri, I. harperi); Koss 1970: 98; Yanoviak and McCafferty 1996: 203; Randolph and McCafferty 1998: 127-128
          (comb.) Siphlurus bicolor (Walker, 1853)
          (syn.) Chirotonetes albomanicatus Needham, 1905
          (syn.) Isonychia albomanicata (Needham, 1905)
          (syn.) Isonychia christina Traver, 1934
          (syn.) Isonychia circe Traver, 1934
          (syn.) Isonychia fattigi Traver, 1934
          (syn.) Isonychia harperi Traver, 1934
          (syn.) Isonychia matilda Traver, 1934
          (syn.) Isonychia pacoleta Traver, 1932
          (syn.) Isonychia sadleri Traver, 1934
        Isonychia rufa McDunnough, 1931c - Randolph and McCafferty 1998: 128-129
        Isonychia sayi Burks, 1953 - WI, IL, IN (Kondratieff and Voshell 1984: 228-231; Randolph and McCafferty 1998: 129), expect in southern Michigan
        Isonychia sicca (Walsh, 1862) Baetis (orig.) - IL, IN, WI (Randolph and McCafferty 1998: 129-130)
          (comb.) Chirotonetes siccus (Walsh, 1862)
          (comb.) Siphlurus siccus (Walsh, 1862)
          (syn.) Chirotonetes mancus (Eaton, 1871)
          (syn.) Isonychia edmundsi Kondratieff & Voshell, 1984
          (syn.) Isonychia manca Eaton, 1871
          (syn.) Isonychia sicca manca Eaton, 1871
          (stat.) Isonychia sicca sicca (Walsh), 1862

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    LEPTOHYPHIDAE Landa and Soldn, 1985

      Tricorythodes Ulmer, 1919
        Tricorythodes allectus (Needham, 1905) Caenis (orig.) - Leonard 1948: 5; Lyman 1956: 574; Koss 1970: 100 (as T. atratus); Randolph and McCafferty 1998: 130
          (syn.) Tricorythodes atratus McDunnough, 1923
          (syn.) Tricorythodes peridius Burks, 1953
          (comb.) Tricorythus allectus (Needham, 1905)
          (syn.) Tricorythus atratus McDunnough, 1923
        Tricorythodes explicatus Eaton, 1892 - Koss 1970: 100 (as T. fallax); Yanoviak and McCafferty 1996: 203 (as T. minutus)
          (syn.) Tricorythodes fallacina McDunnough, 1939
          (syn.) Tricorythodes fallax Traver, 1935
          (syn.) Tricorythodes minutus Traver, 1935
        Tricorythodes mosegus Alba-Tercedor and Flannagan, 1995 - Yanoviak and McCafferty 1996: 203
          (spell.) Tricorythodes mosequs Alba-Tercedor & Flannagan, 1995
        Tricorythodes stygiatus McDunnough, 1931 - Leonard and Leonard 1962: 35

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    LEPTOPHLEBIIDAE Banks, 1900 (Suborder Furcatergalia - forked-gill mayflies)

      Choroterpes Eaton, 1881 (Subfamily Atalophlebiinae Peters, 1980)
        Choroterpes basalis (Banks, 1900) Leptophlebia (orig.) - Lyman 1956: 574; McCafferty 1992b; UMMZ coll.; see also Burian 1995
          (syn.) Choroterpes ferruginea Traver, 1934
          (syn.) Choroterpes fusca Spieth, 1938
          (syn.) Choroterpes hubbelli Berner, 1946

      Habrophlebiodes Ulmer, 1920 (Subfamily Atalophlebiinae Peters, 1980)
        Habrophlebiodes americana (Banks, 1903) Habrophlebia (orig.) Otsego Co., 1972 Leonard collection; however, Randolph and McCafferty (1998: 133) indicate this is a species found primarily in unglaciated areas)
          (syn.) Choroterpes betteni Needham, 1908
          (syn.comb.) Habrophlebiodes betteni (Needham), 1908 (syn.comb.)
          (syn.comb) Leptophlebia betteni (Needham), 1908 (syn.comb.)

      Leptophlebia Westwood, 1840
        Leptophlebia cupida (Say, 1823) Ephemera (orig.) - Needham 1908: 263 (as Blasturus cupidus); Leonard and Leonard 1962: 68, 70; Koss 1970: 99; Randolph and McCafferty 1998: 134
          (comb.) Blasturus cupidus (Say, 1823)
          (comb.) Potamanthus cupidus (Say, 1823)
          (syn.) Baetis ignava Hagen, 1861
          (syn.) Leptophlebia ignava (Hagen, 186)1
          (syn.) Blasturus austrinus Traver, 1932
          (syn.) Leptophlebia austrina (Traver, 1932)
          (syn.) Blasturus collinus Traver, 1932
          (syn.) Leptophlebia collina (Traver, 1932)
          (syn.) Palingenia concinna Walker, 1853
          (syn.) Blasturus concinnus (Walker, 1853)
          (syn.) Leptophlebia concinna (Walker, 1853)
          (syn.) Potamanthus concinnus (Walker, 1853)
          (syn.) Blasturus gravastellus Eaton, 1884
          (syn.) Leptophlebia gravastella (Eaton, 1884)
          (syn.) Cloe quebecensis Provancher, 1876
          (syn.) Ephemerella quebecensis (Provancher, 1876)
          (syn.) Heptagenia quebecensis (Provancher, 1876)
          (syn.) Ephemera hebes Walker, 1853
          (syn.) Leptophlebia hebes (Walker, 1853)
          (syn.) Palingenia pallipes Walker, 1853
          (syn.) Leptophlebia pallipes (Walker, 1853)
        Leptophlebia intermedia (Traver, 1932) Blasturus (orig.)
          (syn.) Blasturus grandis Traver, 1932
          (syn.) Leptophlebia grandis (Traver, 1932)
        Leptophlebia nebulosa (Walker, 1853) Palingenia (orig.) - Lyman 1956: 574; Leonard and Leonard 1962: 67; Koss 1970: 99; Randolph and McCafferty 1998: 135
          (comb.) Potamanthus nebulosus (Walker, 1853)
          (syn.) Blasturus nebulosus (Walker, 1853)
          (syn.) Potamanthus odonatus Walsh, 1862

      Paraleptophlebia Lestage, 1917
        Paraleptophlebia adoptiva (McDunnough, 1929) Leptophlebia - Leonard and Leonard 1962: 73-74; Koss 1970: 99; Yanoviak and McCafferty 1996: 203
        Paraleptophlebia debilis (Walker, 1853) Baetis (orig.) - Leonard and Leonard 1962: 70-71; Koss 1970: 99; Randolph and McCafferty 1998: 136
          (comb.) Leptophlebia debilis (Walker, 1853)
          (id.) Leptophlebia mollis (Hagen, 1861)
          (invalid renam.) Leptophlebia separata Ulmer, 1920
        Paraleptophlebia guttata (McDunnough, 1924a) Leptophlebia (orig.) - Yanoviak and McCafferty 1996: 203
        Paraleptophlebia mollis (Eaton, 1871) Leptophlebia (orig.) - Leonard 1948: 5; Leonard and Leonard 1962: 74-75; Koss 1970: 99; Kovalak 1978; Kohler 1983: 1363; Yanoviak and McCafferty 1996: 203
          (nom. nud.) Cloe mollis Hagen, 1861
          (nom. nud.) Leptophlebia mollis Hagen, 1861
        Paraleptophlebia ontario (McDunnough, 1926a) Leptophlebia (orig.) - IL, IN, OH (Randolph and McCafferty 1998: 139), expect in southern LP (however, see Koss 1970: 99 (as P. praepedita - check record at MSU; and check record in Leonard 1948: 5)
        Paraleptophlebia praepedita (Eaton, 1884) Leptophlebia (orig.) - Leonard and Leonard 1962: 71, 73
        Paraleptophlebia strigula (McDunnough, 1932a) Leptophlebia (orig.) - Leonard and Leonard 1962: 75
        Paraleptophlebia volitans (McDunnough, 1924a) Leptophlebia (orig.) - Yanoviak and McCafferty 1996: 203

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    METRETOPODIDAE

      Metretopus Eaton, 1901
        Metretopus borealis (Eaton, 1871) Heptagenia (orig.) - Leonard and Leonard 1962: 95 (as M. norvegicus); see also Berner 1978: 130-133
          (comb.) Rhithrogena borealis (Eaton, 1871)
          (syn.) Metretopus norvegicus Eaton, 1901

      Siphloplecton Clemens, 1915
        Siphloplecton basale (Walker, 1853) Baetis (orig.) - Lyman 1956: 574; Leonard and Leonard 1962: 96; Koss 1970: 99; Berner 1978: 107-111; Cooper 1992; Randolph and McCafferty 1998: 142
          (comb.) Heptagenia basalis (Walker, 1853)
          (spell.) Siphloplecton basalis (Walker, 1853)
          (syn.) Siphloplecton flexus (Clemens, 1913)
          (syn.) Siphlurus flexus Clemen, 1913s
          (syn.) Siphloplecton signatum Traver, 1932
        Siphloplecton interlineatum (Walsh, 1863) Baetis (orig.) - McCafferty 2009: 365; see also Berner 1978: 122-123
          (hom.) Baetis femorata Walsh, 1862 (B. femorata Say, 1823)
          (renam.) Baetis interlineata Walsh, 1863

    NEOEPHEMERIDAE Needham, Traver & Hsu, 1935

      Neoephemera McDunnough, 1925a
        Neoephemera bicolor McDunnough, 1925a - Berner 1956: 41

    OLIGONEURIIDAE Ulmer, 1914

      Homoeoneuria Eaton, 1881
        Homoeoneuria ammophila (Spieth, 1938) Oligoneuria (orig.) - WI, IL, IN (rare)

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    PALINGENIIDAE Albarda, in Selys-Lonchamps, 1888

      Pentagenia Walsh, 1863
        Pentagenia vittigera (Walsh, 1862) Palingenia (orig.) - UMMZ
          (syn.) Pentagenia quadripunctata Walsh, 1863

    POLYMITARCYIDAE Banks, 1900

      Ephoron Williamson, 1802
        Ephoron album (Say, 1824) Baetis (orig.) - Leonard and Leonard 1962: Koss 1970: 100; Kraft et al. 1978; Herdendorf et al. 1986: 167
          (spell.) Baetis alba Say, 1824
          (comb.) Palingenia alba Say, 1824
          (comb.) Polymitarcys albus Say, 1824
        Ephoron leukon Williamson, 1802 - Koss 1970: 100; Randolph and McCafferty 1998: 145

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    POTAMANTHIDAE Albarda, in Selys-Lonchamps, 1888

      Anthopotamus McCafferty & Bae, 1990
        Anthopotamus myops (Walsh, 1863) Ephemera (orig.) - Koss 1970: 100 (as P. myops), 101 (as P. rufus); Munn and King 1987:119-125
          (syn.) syn species Author, 1934
          (comb.) Potamanthus myops (Walsh, 1863)
          (syn.) Anthopotamus inequalis (Needham, 1909)
          (syn.) Anthopotamus rufous (Argo, 1927)
          (syn.) Potamanthus inequalis Needham, 1909
          (syn.) Potamanthus medius Banks, 1908
          (syn.) Potamanthus rufous Argo, 1927
          (syn.) Potamanthus rufus Argo, 1927
        Anthopotamus verticis (Say, 1839) Baetis (orig.) - Needham 1920: 297
          (syn.) Anthopotamus diaphanus (Needham, 1907)
          (syn.) Potamanthus diaphanus Needham, 1907
          (syn.) Anthopotamus walkeri (Ide, 1935)
          (syn.) Ecdyurus verticis Walsh, 1862
          (comb.) Heptagenia verticis (Say, 1839)
          (syn.) Potamanthus betteni Morgan, 1913
          (syn.) Potamanthus flaveola (Walsh, 1862)
          (comb.) Potamanthus verticis (Say, 1839)
          (syn.) Potamanthus walkeri Ide, 1935

    PSEUDIRONIDAE

      Pseudiron McDunnough, 1931a
        Pseudiron centralis McDunnough, 1931a - McCafferty 1992a
          (syn.) Pseudiron meridionalis Traver, 1935

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    SIPHLONURIDAE Ulmer, 1920

      Parameletus Bengtsson, 1908
        Parameletus croesus (McDunnough, 1923 as Siphlonuroides) - MN, WI (Burian 2017), expect in the w. UP

      Siphlonurus Eaton, 1868
        Siphlonurus alternatus (Say, 1824) Baetis (orig.) - Leonard 1948: 5; Leonard and Leonard 1962: 77; Koss 1970: 98; Randolph and McCafferty 1998: 149
          (syn.) Baetis annulata Walker, 1853
          (syn.spell.) Baetis annulatus Walker, 1853
          (syn.) Baetis femorata Provancher, 1876
          (syn.) Siphlurella alternata Say, 1824
          (syn.spell.) Siphlurus alternatus Say, 1824
        Siphlonurus mirus (Eaton, 1885a) Siphlurus - ON (McCafferty et al. 2008: 447)
        Siphlonurus quebecensis (Provancher, 1878) Siphlurus (orig.) - Leonard 1948: 5; Leonard and Leonard 1962: 78; Koss 1970: 98; Randolph and McCafferty 1998: 150
          (syn.) Siphlonurus triangularis Clemens, 1915
        Siphlonurus phyliis McDunnough, 1923 - Dewalt & South 2015: 145 (locations 47.92155N -89.14587W, 47.93499N -89.17524W, 47.93854N -89.15144W, 47.91959N -89.17672W, 47.91965N -89.15921W, 48.12657N -88.53880W)
        Siphlonurus rapidus McDunnough, 1924c - Leonard 1948: 5; Lyman 1956: 574; Leonard and Leonard 1962: 76; Koss 1970: 98; Randolph and McCafferty 1998: 151
        Siphlonurus typicus (Eaton, 1885a) Siphlurus orig. - Koss 1970: 98; Randolph and McCafferty 1998: 151
          (syn.) Siphlonurus berenice McDunnough, 1923
          (syn.spell.) Siphlonurus bernice McDunnough, 1923
          (syn.) Siphlonurus novangliae McDunnough, 1924c
References - Distribution
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    Allen RK, Edmunds GF. 1962b. A revision of the genus Ephemerella (Ephemeroptera, Ephemerellidae) IV. The subgenus Dannella in North America. Journal of Kansas Entomological Society 35(3):333-338.
    Allen RK, Edmunds GF. 1963a. A revision of the genus Ephemerella (Ephemeroptera, Ephemerellidae) VI. The subgenus Serratella in North America. Annals of the Entomological Society of America 56(5): 583-600.
    Allen RK, Edmunds GF. 1963b. A revision of the genus Ephemerella (Ephemeroptera, Ephemerellidae) VII. The subgenus Eurylophella in North America. The Canadian Entomologist 95(6):597-623.
    Allen RK, Edmunds GF. 1965. A revision of the genus Ephemerella (Ephemeroptera, Ephemerellidae) VIII. The subgenus Ephemerella in North America. Miscellaneous Publications of the Entomological Society of America 4(6):243-282.
    Banks N. 1910. Notes on our eastern species of the May-fly genus Heptagenia. The Canadian Entomologist 21: 197-202.
    Bednarik AF, McCafferty WP. 1979. Biosystematic revision of the genus Stenonema (Ephemeroptera: Heptageniidae). Canadian Bulletin of Fisheries and Aquatic Sciences 201: vi, 1-73.
    Berner L. 1978. A review of the mayfly family Metretopodidae. Transactions of the American Entomological Society 104(1): 91-137.
    Burian SK. 1995. Taxonomy of the eastern Nearctic species of Choroterpes Eaton (Ephemeroptera: Leptophlebiidae), pp. 433-454 in Corkum LD, Ciborowski JJH (eds.). Current Directions in Research on Ephemeroptera. Canadian Scholars' Press Inc., Toronto, Ontario, Canada.
    Burian SK. 2017. The taxonomy of the North American species of Parameletus Bengtsson, 1908 (Ephemeroptera: Siphlonuridae), with keys to nymphs and male imagos. Canadian Journal of Arthropod Identification 33: 1-76.
    Cooper JC. 1993. Life history and ecology of Siphloplecton basale (Walker) (Metretopodidae: Ephemeroptera) in three Michigan trout streams. M.S. thesis, Central Michigan University, Mount Pleasant, Michigan. 206 p.
    DeWalt RE, South EJ. 2015. Ephemeroptera, Plecoptera, and Trichoptera on Isle Royale National Park, USA, compared to mainland species pool and size distribution. ZooKeys 532: 137-158.
    Duffy WG, Batterson TR, McNabb. 1987. The St. Marys River, Michigan: an ecological profile. U.S. Fish and Wildlife Service Biological Report 85 (7.10):1-138.
    Funk DH, Sweeney BW. 1994. The larvae of eastern North American Eurylophella Tiensuu (Ephemeroptera: Ephemerellidae). Transactions of the American Entomological Society 120(3): 209-286.
    Gorski MR, Fox ADE, McQueen JI, Jacobus LM. 2015. Plauditus cestus (Provonsha & McCafferty, 1982) (Insecta: Ephemeroptera: Baetidae): New records from Virginia and the Northwest Territories, with notes on color variation. Check List 11(1, Article 1498):13.
    Jacobus LM. 2013. A new junior synonym for the Holarctic species Ametropus fragilis Albarda, 1878 (Insecta: Ephemeroptera: Ametropodidae). Proceedings of the Indiana Academy of Science 122(1):1819.
    Jacobus LM , McCafferty WP. 2004. Revisionary contributions to the genus Drunella (Ephemeroptera: Ephemerellidae). Journal of the New York Entomological Society 112(2-3):127-147.
    Jacobus LM , Wiersema NA. 2014. The genera Anafroptilum Kluge, 2011 and Neocloeon Traver, 1932, reinstated status, in North America, with remarks about the global composition of Centroptilum Eaton, 1869 (Ephemeroptera: Baetidae). Zootaxa 3814(3):385391.
    Herdendorf CE, Raphael CN, Jaworski E. 1986. The ecology of Lake St. Clair wetlands: a community profile. U.S. Fish and Wildlife Service Biological Report 85 (7.7).
    Hill MA, Pfeiffer J, Jacobus LM. 2010. A new genus and new species of Baetidae (Ephemeroptera) from lakes and reservoirs in eastern North America. Zootaxa 2481: 61-68.
    Hunt BP. 1953 The life history and economic importance of a burrowing mayfly, Hexagenia limbata, in southern Michigan lakes. Bulletin of the Institute for Fisheries Research 4:1-151.
    Koss RW. 1970. A list of the mayflies (Ephemeroptera) in the Michigan State University entomology museum. The Michigan Entomologist 3(4): 98-101.
    Kovalak WP. 1978. Diel changes in stream benthos density on stones and artificial substrates. Hydrobiologia 58(1): 7-16.
    Kraft KJ, Johnson K, Sypniewski RH. 1978. Mating flights of Ephoron album (Ephemeroptera: Polymitarcidae) in Michigan. The Great Lakes Entomologist 11(2):85-88.
    Leonard JW. 1948. The seasonal occurrence of mayflies in a Michigan stream. Institute for Fisheries Research, Division of Fisheries, Michigan Department of Conservation. Report No. 1165, March 9, 1948. 8 pp.
    Leonard JW, Leonard FA. 1962. Mayflies of Michigan trout streams. Cranbrook Institute of Science, Bloomfield Hills, Michigan, USA. x + 139 p.
    Lewis PA. 1974. Taxonomy and ecology of Stenonema mayflies (Heptageniidae: Ephemeroptera). National Environmental Research Center Office of Research and Development. U.S. EPA. Cincinnati, Ohio. EP-670/4-74-006. 80 p.
    Lyman FE. 1945. Reactions of certain nymphs of Stenonema (Ephemeroptera) to light as related to habitat preference. Annals of the Entomological Society of America 38(2):234-236.
    Lyman FE. 1955. Seasonal distribution and life cycles of Ephemeroptera. Annals Entomological Society of America 41: 380-391.
    Lyman FE. 1956. Environmental factors affecting distribution of mayfly nymphs in Douglas Lake, Michigan. Ecology 37(3): 568-576.
    McCafferty WP. 1992a. The sand-dwelling predatory mayfly Pseudiron centralis in Michigan (Ephemeroptera: Pseudironidae). The Great Lakes Entomologist 25(2): 133-134.
    McCafferty WP. 1992b. New larval descriptions and comparisons of North American Choroterpes (Ephemeroptera: Leptophlebiidae). The Great Lakes Entomologist 25(2): 71-78.
    McCafferty WP. 2009. New state and provincial North American records for 100 Ephemeroptera species. Transactions of the American Entomological Society 135(3):353-368.
    McCaffery WP, Lenat DR. 2003. A new Nearctic Paracloeodes (Ephemeroptera: Baetidae). Entomological News 114(1):33-36.
    McCafferty WP, Jacobus LM, Webb JM, Meyer MD. 2008. Insecta, Ephemeroptera: Range extensions and new records for Ontario and Canada. Check List 4(4): 445448.
    Munn MD, King RH. 1987. Life history of Potamanthus myops (Walsh)(Ephemeroptera: Potamanthidae) in a Central Michigan stream. American Midland Naturalist 117(1): 119-125.
    Needham JG. 1908. Notes on the aquatic insects of Walnut Lake. Report of the Board of Geological Survey of Michigan for 1907 on the Biology of Walnut Lake by T. L. Hankinson and others: 259-263.
    Needham JB. 1920. Burrowing mayflies from our lakes and streams. Bulletin of the Bureau of Fisheries 36: 269-304.
    Pescador ML, Berner L. 1981. The mayfly family Baetiscidae (Ephemeroptera). Part II. Biosystematics of the genus Baetisca. Transactions of the American Entomological Society 107(3):163-228.
    Randolph RP, McCafferty WP. 1998. Diversity and distribution of the mayflies (Ephemeroptera) of Illinois, Indiana, Kentucky, Michigan, Ohio, and Wisconsin. Ohio Biological Survey Bulletin 13(1): vii, 1-188.
    Randolph RP, McCafferty WP, Zaranko D, Jacobus LM, Webb JM. 2002. New Canadian records of Baetidae (Ephemeroptera) and adjustments to North American Cloeon. Entomological News 113(5):306-308.
    Spieth HT. 1941. Taxonomic studies on the Ephemeroptera. II. The genus Hexagenia. The American Midland Naturalist 26(2): 233-280.
    Steven JC, Hilsenhoff WL. 1979. Ametropus neavei (Ephemeroptera: Ametropodidae) in the upper peninsula of Michigan. The Great Lakes Entomologist 12(4): 226.
    Sun L, McCafferty WP. 2008. Cladistics, classification and identification of the brachycercine mayflies (Insecta: Ephemeroptera: Caenidae). Zootaxa 1801: 1239
    Waltz RD, Munro JB. 1996. Procloeon viridocularis (Ephemeroptera: Baetidae) from Michigan and Pennsylvania, new range extensions with comments on the species. Entomological News 107(3):166-168
    Waltz RD, Burian SK. 2007. Ephemeroptera, pp. 181-236 in Merritt RW, Cummins KW, Berg MB (eds.) An Introduction to the Aquatic Insects of North America, Fourth Edition. Kendall/Hunt Publishing Company, Dubuque, Iowa.
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References - Taxonomic

Alba-Tercedor J, Flannagan J. 1995. Two new Canadian species of the genus Tricorythodes Ulmer, with additional studies on other North American species (Insecta, Ephemeroptera: Leptohyphidae). Canadian Journal of Zoology 73: 1588-1598.
Albarda H. 1878. Descriptions of three new European Ephemeridae. Entomologist's Monthly Magazine 15: 128-130.
Allen RK. 1968. New species and records of Ephemerella (Ephemerella) in western North America (Ephemeroptera: Ephemerellidae). Journal of the Kansas Entomological Society 41(4):557-567.
Allen RK, Edmunds GF. 1962a. A revision of the genus Ephemerella (Ephemeroptera, Ephemerellidae) V. The subgenus Drunella in North America. Miscellaneous Publications of the Entomological Society of America 3(5):147-179.
Allen RK, Edmunds GF. 1962b. A revision of the genus Ephemerella (Ephemeroptera, Ephemerellidae) IV. The subgenus Dannella in North America. Journal of Kansas Entomological Society 35(3):333-338.
Allen RK, Edmunds GF. 1963a. A revision of the genus Ephemerella (Ephemeroptera, Ephemerellidae) VI. The subgenus Serratella in North America. Annals of the Entomological Society of America 56(5): 583-600.
Allen RK, Edmunds GF. 1963b. A revision of the genus Ephemerella (Ephemeroptera, Ephemerellidae) VII. The subgenus Eurylophella in North America. The Canadian Entomologist 95(6):597-623.
Allen RK, Edmunds GF. 1965. A revision of the genus Ephemerella (Ephemeroptera, Ephemerellidae) VIII. The subgenus Ephemerella in North America. Miscellaneous Publications of the Entomological Society of America 4(6):243-282.
Argo VN. 1927. The North American species of the genus Potamanthus, with a description of a new species. Journal of the New York Entomological Society 35(4): 319-329.
Balthasar V. 1937. Arthropleidae, eine neue Familie der Ephemeropteren. Zoologischer Anzeiger 120(9-10): 204-230.
Banks N. 1900. New genera and species of Nearctic Neuropteroid Insects. Transactions of the American Entomological Society 26: 239-259.
Banks N. 1903. A new species of Habrophlebia. Entomological News 14: 235.
Banks N. 1907. A new mayfly of the genus Caenis. Entomological News 18: 13-14.
Banks N. 1908. Neuropteroid insects - notes and descriptions. Transactions of the American Entomological Society 34: 255-267.
Banks N. 1914. New neuropteroid insects, native and exotic. Proceedings of the Academy of Natural Sciences of Philadelphia 66: 608-632, pl. 28.
Banks N. 1924. Descriptions of new neruopteroid insects. Bulletin of the Museum of Comparative Zoology 65(12): 419-455, pl. 1-4.
Bednarik AF. 1979. Subgeneric classification of Stenonema (Ephemeroptera: Heptageniidae). Journal of the Georgia Entomological Society 14(3): 190-191.
Bednarik AF, McCafferty WP. 1979. Biosystematic revision of the genus Stenonema (Ephemeroptera: Heptageniidae). Canadian Bulletin of Fisheries and Aquatic Sciences 201: vi, 1-73.
Bengtsson S. 1909. Beiträge zur Kenntnis der paläarktischen Ephemeriden. Lunds Universitets Årsskrift, N. F., Afd. 2, 5(4): 1-19.
Bengtsson S. 1913. Undersökningar öfver äggen hos Ephemeriderna. Entomologisk Tidskrift 34(2-4): 271-320, pl. 1-3.
Bengtsson S. 1912. Neue Ephemeriden aus Schweden. Entomologisk Tidskrift 33(1-2): 107-117.
Bengtsson S. 1914. Bemerkungen Über die nordischen Arten der Gattung Cloëon Leach. Entomologisk Tidskrift 35(3-4): 210-220.
Bengtsson S. 1915. Eine Namensänderung. Entomologisk Tidskrift 36(1): 34.
Bengtsson S. 1936. Eine neue Cloeon-Art aus Danmark, Cl. Schoenemundi n.sp. Opuscula Entomologica 1(1): 35.
Berner L. 1940. Baetine mayflies from Florida (Ephemeroptera). Florida Entomologist 23: 33-45, 49-62.
Berner L. 1946. New species of Florida mayflies (Ephemeroptera). Florida Entomologist 28:60-82.
Berner L. 1978. A review of the mayfly family Metretopodidae. Transactions of the American Entomological Society 104(1): 91-137.
Berner L, Allen RK. 1961. Southeastern species of the mayfly subgenus Serratella (Ephemerella: Ephemerellidae). Florida Entomologist 44: 149-158.
Burian SK. 1995. Taxonomy of the eastern Nearctic species of Choroterpes Eaton (Ephemeroptera: Leptophlebiidae), pp. 433-454 in Corkum LD, Ciborowski JJH (eds.). Current Directions in Research on Ephemeroptera. Canadian Scholars' Press Inc., Toronto, Ontario, Canada.
Burian SK. 2017. The taxonomy of the North American species of Parameletus Bengtsson, 1908 (Ephemeroptera: Siphlonuridae), with keys to nymphs and male imagos. Canadian Journal of Arthropod Identification 33: 1-76.
Burks BD. 1946. New heptagenine mayflies. Annals of the Entomological Society of America 39(4): 607-615.
Burks BD. 1949. New species of Ephemerella from Illinois (Ephemeroptera). The Canadian Entomologist 79: 232-236.
Burks BD. 1953. The mayflies, or Ephemeroptera, of Illinois. Bulletin of the Illinois Natural History Survey 26(1): 1-216.
Carle FL. 1978. A new species of Ameletus (Ephemeroptera: Siphlonuridae) from western Virginia. Annals of the Entomological Society of America 71(4): 581-584.
Clemens WA. 1913. New species and new life histories of Ephemeridae or mayflies. The Canadian Entomologist 45: 246-262, 329-341, pl. 5-7.
Clemens WA. 1915. Mayflies of the Siphlonurus group. The Canadian Entomologist 47:245-260, pl. 9-11.
Daggy RH. 1945. New species and previously undescribed naiads of some Minnesota mayflies (Ephemeroptera). Annals of the Entomological Society of America 38(3): 373-396.
Dodds GS. 1923. Mayflies from Colorado. Descriptions of certain species and notes on others. Transaction of the American Entomological Society 49: 93-114, pl. 8-9.
Day WC. 1955. New genera of mayflies from California (Ephemeroptera). Pan-Pacific Entomologist 31(3): 121-137.
Eaton AE. 1868. An outline of a re-arrangement of the genera of Ephemeridae. Entomologist's Monthly Magazine 5: 82-91.
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Eaton AE. 1883. pp. 1-76 in A revisional monograph of recent Ephemeridae or mayflies, Part I. (1883-1888). Transactions of the Linnean Society of London, Second Series, Zoology 3: 1-352, 65 pl.
Eaton AE. 1884. pp. 77-152 in A revisional monograph of recent Ephemeridae or mayflies, Part II. (1883-1888). Transactions of the Linnean Society of London, Second Series, Zoology 3: 1-352, 65 pl.
Eaton AE. 1885a. pp. 153-230 in A revisional monograph of recent Ephemeridae or mayflies, Part III. (1883-1888). Transactions of the Linnean Society of London, Second Series, Zoology 3: 1-352, 65 pl.
Eaton AE. 1885b. pp. 229-282 in A revisional monograph of recent Ephemeridae or mayflies, Part III. (1883-1888). Transactions of the Linnean Society of London, Second Series, Zoology 3: 1-352, 65 pl.
Eaton AE. 1892. Fam. Ephemeridae. Biologia Centrali-Americana 38: 1-16, 1 pl.
Eaton AE. 1901. Ephemeridae collected by Herr E. Strand in South and Arctic Norway. Entomologist's Monthly Magazine 37: 252-255.
Eaton AE. 1908. In P. Esben-Petersen, Bidrag til en Fortegnelse over arktisk Norges Neuropterfauna. (With description of a new species of Ephemeridae by Rev. A. E. Eaton). Tromsø Museums Aarshefter (1902) 25:119-153.
Edmunds GF Jr. 1959. Subgeneric groups within the mayfly genus Ephemerella (Ephemeroptera: Ephemerellidae). Annals of the Entomological Society of America 52(5): 543-547.
Edmunds GF Jr. 1971. A new name for a subgeneric homonym in Ephemerella (Ephemeroptera: Ephemerellidae). Proceedings of the Entomological Society of Washington 73(2): 152.
Edmunds GF Jr, Allen RK. 1957. A new species of Baetis from Oregon (Ephemeroptera: Baetidae). Journal of the Kansas Entomological Society 30: 57-58.
Edmunds GF Jr, Jensen SL. 1974. A new genus and subfamily of North American Heptageniidae (Ephemeroptera). Proceedings of the Entomological Society of Washington 76(4): 495-497.
Edmunds GF Jr., Traver JR. 1959. The classification of the Ephemeroptera. I. Ephemeroidea: Behningiidae. Annals of the Entomological Society of America 52(1): 43-51.
Edmunds GF Jr. 1963. In Edmunds GF Jr, Allen RK, Peters WL, An annotated key to the nymphs of the families and subfamilies of mayflies (Ephemeroptera). University of Utah Biological Series 13(1): 1-49.
Esben-Petersen P. 1909. New Ephemeridae from Denmark, Arctic Norway and the Argentine Republic. Deutsche Entomologische Zeitschrift 1909: 551-556.
Flowers RW. 1980. Two new genera of Nearctic Heptageniidae (Ephemeroptera). Florida Entomologist 63: 296-307.
Funk DH, Sweeney BW. 1994. The larvae of eastern North American Eurylophella Tiensuu (Ephemeroptera: Ephemerellidae). Transactions of the American Entomological Society 120(3): 209-286.
Gorski MR, Fox ADE, McQueen JI, Jacobus LM. 2015. Plauditus cestus (Provonsha & McCafferty, 1982) (Insecta: Ephemeroptera: Baetidae): New records from Virginia and the Northwest Territories, with notes on color variation. Check List 11(1, Article 1498):13.
Gose K. 1980. Nihon san kagero-rui. [The mayflies of Japanese. Key to families, genera and species.] (in Japanese). Aquabiology (Nara) 2(1): 76-79; 2(2): 122-123; 2(3): 211-215; 2(4): 286-288; 2(5): 366-368; 2(6): 454-457.
Hagen HA. 1861. Synopsis of the Neuroptera of North America. With a list of the South American species. Smithosnian Miscellaneous Collections, Washington. pp 1-347.
Hill MA, Pfeiffer J, Jacobus LM. 2010. A new genus and new species of Baetidae (Ephemeroptera) from lakes and reservoirs in eastern North America. Zootaxa 2481: 61-68.
Hubbard MD. 1974. Some specific homonyms in mayflies (Ephemeroptera). Journal of the Kansas Entomological Society 47(3):357-359.
Ide FP. 1930. Contibution to the biology of Ontario mayflies with descriptions of new species. The Canadian Entomologist 62: 204-213, 218-231.
Ide FP. 1935. Life history notes on Ephoron, Potamanthus, Leptophlebia and Blasturus with descriptions (Ephemeroptera). The Canadian Entomologist 67(6): 113-125.
Ide FP. 1937. Descriptions of eastern North American species of baetine mayflies with particular reference to the nymphal stages. The Canadian Entomologist 69:219-231.
Ide FP. 1954. The nymph of Rhithrogena impersonata (Ephemerida) and a new closely related species from the same locality in southern Ontario. The Canadian Entomologist 86(8): 348-356.
Imanishi K. 1940. Mayflies from Manchuria, Lower Mongolia, and Korea. Report of the Limnological Society of Kwantung and Manchoukuo. Darien. Pages 169-263. (In Japanese)
Jacob U. 1986. In Jacob U, Glazaczow A. 1986. Pseudocentroptiloides, a new baetid genus of Palaearctic and Oriental disbtribution [sic] (Ephemeroptera). Aquatic Insects (1986) 8(4): 197-206.
Jacobus LM. 2013. A new junior synonym for the Holarctic species Ametropus fragilis Albarda, 1878 (Insecta: Ephemeroptera: Ametropodidae). Proceedings of the Indiana Academy of Science 122(1):1819.
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Page edits
    November 03, 2017
    • Revised section on Parmeletus, to reflect information in Burian 2017. A regional record for P. chelifer Bengtsson, 1908, based on a specimen in Wisconsin, was misidentified and has been determined to be P. croesus (McDunnough, 1923), which also occurs in Minnesota. P. chelifer is now recognized to occur in the far north and northern Europe.
    March 10, 2017
    • Reorganized reference sections, splitting them into references pertaining to sources documenting the occurence of a species in Michigan, and another for works that reference authorship of species cited.
    March 06, 2015
    • Baetis bundyae and Siphlonurus phyllis recorded from Isle Royale, per DeWalt & South 2015..
    January 12, 2015
    • Pladitus cestus added as likely species to be found in Michigan, per Gorski et al. 2015
    July 24, 2014
    • Corrected authorship error for Cercobrachys fox, C. lilliei, and C. winnegago. Thanks to Luke Jacobus for noticing this error.
    June 24, 2014
    • Centroptilum does not occur in North America, but our species have been reassigned to Anafroptilum and Neocloeon (alamance and triangulifer), per Jacobus & Wiersema 2014.
    April 16, 2014
    • Ametropus neavei a junior synonym of Ametropus fragilis per Jacobus 2014.
    January 06, 2013
    • Removed Maccaffertium ithaca as a Michigan record (based on Leonard and Leonard 1962: 108) per Bednarik and McCafferty (1979: 23), who indicate Michigan and Minnesota records are likely misidentifications, as this species is essentially Appalachian in distribution.
    December 27, 2012
    • Removed Raptoheptagenia cruentata as a possible Michigan species; its range in IL, IN and OH apparently is restricted to Mississippi-draining systems; a widespread species (also known from AB and SK), it probably doesn't occur in Michigan
    • Pseudocloeon dardanum, P. ephippitum, P. frondalis, and P. propinquum moved back to Labiobaetis
    • Removed Callibaetis ferrugineus hageni, as its distribution is western North America
    November 21, 2012
    • Revised html coding, reducing file size
    November 01, 2012
    • Beginning to keep track of changes to the web site pages
    • Removed individual pages for families into one long list, families accessed from left menu via anchor redirects within page
    • Updated synonomies, based on Mayfly Central and other publications
    • Pentagenia vittigera (Walsh, 1862) moved from Ephemeridae to Palingeniidae (see McCafferty and Edmunds 1976)
    • Added Cloeon dipterum (as Cloeon cognatum) as new Michigan record; specimen collected by K.Perez, S.McCubbin, J.Cossey, A.Borisenko - Oceana Co. (43.4851N, -86.3603); data from supplementary spreadsheet from publisher of Webb et al. 2012).
Page created: October 29, 2002 - Last updated: November 3, 2017 (EB)