Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

Toggle Menu

Aquatic Coleoptera (Beetles) of Michigan

Introduction

NOTE: This page is undergoing a revision, and a new version should appear in late August 2017. To reduce length, a number of family species lists will appear on separate pages. Each family will have their own separate reference sections for distribution and taxonomy, and numerous omissions and errors in citations will be corrected.

Certain groups of aquatic beetles have been well surveyed in our state, and several systematic monographs have appeared for several important families (e.g., Dytiscidae) that occur in our state. Still, the enormous number of Coleopteran species that may occur in our state means that to fashion completed species lists for all the families listed to the left will require additional work. More information regarding aquatic beetles and Coleoptera in genera can be found at the Coleoptera Home Page.

Genera are donoted in bold green; species denoted in bold blue have been recorded in Michigan; other species denoted in bold are likely to occur based on their known distribution. You can use the Find function in your web browser to locate families, genera, and species. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated.

Adephaga Schellenberg, 1806

    DYTISCIDAE Leach, 1815: 68 (Predaceous Diving Beetles) (separate page)

     
     

    GYRINIDAE Latreille, 1802 (Whirligig Beetles)

    [under contruction] Gyrinids, or whirligig beetles, are aquatic beetles that are familiar to most everyone by characteristic circular movements on the water surface. Medium-sized adults are shiny and usually brown-bronze to black in color. Larvae and adults are aquatic, with adults being scavengers and larvae predators that use ttheir mandibles to inject poison to subdue prey. Overwinter and females oviposit under water.

    Adult Dineutus sp. (Copyright © 2006 Ethan Bright)
      Dineutus MacLeay, 1825*: 30
        Dineutus assimilis (Kirby, 1837*: 78 as Cyclinus) - UMMZ, NMNH (web page database); Gustafson & Miller 2015*: 123
        Dineutus discolor Aubé, 1838*: 784 - NMNH (web page database); Gustafson & Miller 2015: 123
          (syn.) Cyclous labratus Melsheimer, 1846*: 9
        Dineutus emarginatus (Say, 1825*: 108 as Gyrinus) - UMMZ; see also Hilsehoff 1990: 83, 85; Gustafson & Miller 2015: 124
        Dineutus hornii Roberts, 1895*: 282 - UMMZ, NMNH (web page database); Gustafson & Miller 2015*: 123
        Dineutus nigrior Roberts, 1895*: 280 - UMMZ; Gustafson & Miller 2015: 123

      Gyrinus Geoffroy, 1762
        Gyrinus aeneolus LeConte, 1868*: 368, 370 - UMMZ, NMNH (web page database); Oygur and Wolfe 1990: 59-60, 68
        Gyrinus aeratus Stephens, 1835*: 395- ON, NY (expect in UP, n. LP) (Oygur and Wolfe 1990: 38-39, 72)
          (syn.) Gyrinus aeneus Thomson, 1860*: 116
          (syn.) Gyrinus thomsoni Zaitsev, 1907*: 122
          (syn.) Gyrinus edwardsi Sharp, 1914*: 131, 137
          (syn.) Gyrinus instabilis Fall, 1931*: 155
        Gyrinus affinis Aubé, 1838*: 669- UMMZ, NMNH (web page database); Oygur and Wolfe 1990: 30-33, 52
          (syn.) Gyrinus canadensis Régimbart, 1883*: 159
        Gyrinus analis Say, 1825*: 108 - UMMZ, NMNH (web page database); Oygur and Wolfe 1990: 62-63,67
        Gyrinus aquiris LeConte, 1868*: 368, 371 - UMMZ, NMNH (web page database); Oygur and Wolfe 1990: 66, 68-70
        Gyrinus bifarius Fall, 1922*: 276, 286 - UMMZ; Oygur and Wolfe 1990: 40-42, 70
          (syn.) Gyrinus punctellus Ochs, 1949*: 258
        Gyrinus borealis Aubé, 1838*: 692 - Hilsenhoff 1990: 88; Oygur and Wolfe 1990: 45-46, 50; UMMZ
          (syn.) Gyrinus corpulentus Régimbart, 1883*: 178
        Gyrinus conifinis LeConte, 1868*: 368, 370 - UMMZ, record at Albertson College of Idaho Collection; Oygur and Wolfe 1990: 54-55, 70
        Gyrinus dubius Wallis, 1926*: 93 - northern, may occur on Isle Royale (Oygur and Wolfe 1990: 65, 73)
        Gyrinus dichrous LeConte, 1868*: 368, 371 - Oygur and Wolfe 1990: 61-62, 71; UMMZ
        Gyrinus fraterunus Couper, 1865*: 60 - Mackinaw Co. (Merritt et al. 2002: 155)
        Gyrinus gehringi Chamberlain, 1929*: 247- Oygur and Wolfe 1990: 66-67
        Gyrinus gibber LeConte, 1868*: 370, 372 - Oygur and Wolfe 1990: 48-49, 51; UMMZ
          (syn.) Gyrinus frosti Fall, 1922*: 278, 303
          (syn.) Gyrinus floridensis Ochs, 1929*: 123
        Gyrinus impressicollis Kirby, 1837*: 79 - Oygur and Wolfe 1990: 36, 72
        Gyrinus latilimbus Fall, 1922*: 368, 370 - Oygur and Wolfe 1990: 53-54, 71; UMMZ
        Gyrinus lecontei Fall, 1922*: 291 - UMMZ; Oygur and Wolfe 1990: 66-68
        Gyrinus maculiventris LeConte, 1868*: 368, 371 - Oygur and Wolfe 1990: 64, 67; UMMZ
        Gyrinus marginellus Fall, 1922*: 276, 283 - Oygur and Wolfe 1990: 46, 48, 69; UMMZ
        Gyrinus minutus Fabricius, 1798*: 65 - Oygur and Wolfe 1990: 25-27, 30-31 (however, this may be G. cavatus Atton, 1990 - check article in Can Ent. that O & W were unaware of); UMMZ
          (syn.) Gyrinus kirbyi Marsham, 1802*: 100
        Gyrinus pectoralis LeConte, 1868*: 370, 372 - Oygur and Wolfe 1990: 24, 72; Merritt et al. 2002: 155; UMMZ
        Gyrinus pernitidus LeConte, 1868*: 396, 372 - NY, WI (Oygur and Wolfe 1990: 51, 56-57)
        Gyrinus piceolus Blatchely, 1910*: 238, 240 - IN (known only from Indiana, but might occur in sw. LP) (Oygur and Wolfe 1990: 49, 51)
        Gyrinus picipes Aubé, 1838*: 694 - ON, expect in w. UP, Isle Royale (Oygur and Wolfe 1990: 64-65, 73)
        Gyrinus pugionis Fall, 1922*: 277, 296 - Oygur and Wolfe 1990: 46, 50; UMMZ
        Gyrinus sayi Aubé, 1838*: 698 - Oygur and Wolfe 1990: 65-66, 73; UMMZ
          (syn.) Gyrinus lugens LeConte, 1868*: 396, 372
          (syn.) Gyrinus falli Notman, 1919*: 131
        Gyrinus ventralis Kirby, 1837*: 80 - Oygur and Wolfe 1990: 49-51; Merritt et al. 2002: 155; UMMZ
          (syn.) Gyrinus hatchi Wallis, 1926*: 93
        Gyrinus wallisi Fall, 1922*: 278 - MN, ON, expect in UP (Oygur and Wolfe 1990: 36-37, 72)
        Gyrinus woodruffi Fall, 1922*: 276, 282 - UMMZ

      References - Taxonomic and Distribution

        Aubé C. 1836–1838. Tome Cinquième (V). Hydrocanthares. Méquignon-Marvis Père et fils, Paris, xi + 415 pp. + 446 pls. [411–464: 1836, 1865–1224: 1837, 1225–1416: 1838.]
        Blatchley WS. 1910. An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of the Rhynchophoral known to occur in Indiana with bibliography and descriptions of new species. Bulletin no. 1 Indiana Department of Geology and Natural Resources. Indianapolis, Indiana, 1386 pp., illus. Reprinted by Nature Publishing Co. in 1910.
        Chamberlain KF. 1929. A new species of Gyrinus from Northern New Hampshire. Bulletin of the Brooklyn Entomological Society 24: 247-249.
        Couper W. 1865. Description of new species of Canadian Coleoptera. Canadian Naturalist 2(2): 60-63.
        Fabricius JC. 1798. Entomologiae Systematica. Supplement. Proft and Storch, Hafniae. [2] + 572 + 52 pp. (PDF)
        Fall HC. 1922. The North American species of Gyrinus (Coleoptera). Transactions of the American Entomological Society 47: 269-306 + pl. 16.
        Fall HC. 1931. A new Gyrinus from Alaska with reference to other recently described species (Coleoptera). Pan-Pacific Entomologist 7(4): 154-156.
        Geoffroy E-L. 1762. Histoire abregée des insectes qui se trouvent aux environs de Paris, dans laquelle ces animaux sont rangés suivant un ordre méthodique. Chez Durand, Paris, 523 pp. + 510 pl.
        Gustafson GT, Miller KB. 2015. The New World whirligig beetles of the genus Dineutus Macleay, 1825 (Coleoptera, Gyrinidae, Gyrininae, Dineutini). ZooKeys 476: 1–135. doi: 10.3897/zookeys.476.8630.
        Hilsenhoff WL. 1990. Gyrinidae of Wisconsin, with a key to adults of both sexes and notes on distribution and habitat. The Great Lakes Entomologist 23(2):77-91.
        Kirby W. 1837. The Insects. in Richardson J, Fauna boreali-americana, or, The zoology of the northern parts of British America: containing descriptions of the objects of natural history collected on the late northern land expeditions, under command of Captain Sir John Franklin, R.N., Part 4. London. xxxix + 249 pp. VIEW
        LeConte JL. 1868. The Gyrinidae of America, north of Mexico. Proceedings of the Academy of Natural Sciences of Philadelphia 20: 365–373.
        Macleay WS. 1825. Annulosa javanica, or an attempt to illustrate the natural affinities and analogies of the insects collected in Java by Thomas Horsfiled and deposited by him in the museum of the Honourable East-India Company. Kingsbury, Parbury and Allen, London, xii + 50 pp. (PDF)
        Marsham T. 1802. Entomologia Britannica, sistens Insecta Britanniae indigena, secundum methodum Linnaeanum disposita. Londini 1: 1-548.
        Melsheimer FE. 1846. Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 2: 26–43.
        Merritt RW, Benbow ME, Hudson PL. 2002. Wetland macroinvertebrates of Prentiss Bay, Lake Huron, Michigan: diversity and function group comparison. The Great Lakes Entomologist 35(2): 149-160.
        Notman H. 1919. Coleoptera collected at Mooers, Clinton Co., N.Y. September 9-13, 1918 with descriptions of new species. Bulletin of the Brooklyn Entomological Society 129: 131-132.
        Ochs G. 1929. Notes upon some Gyrinidae in the Carnegie Museum with descriptions of new species. Annals of the Carnegie Museum 9: 123-134.
        Ochs G. 1949. A revision of the Gyrinoidea of Central America (Col). Revista de Entomologia (Brazil) 20(1-3): 253–300.
        Régimbart M. 1883. Essai monographique de la famille des Gyrinidae. Annales de la Société Entomologique de France (6)3: 121–190.
        Roberts CH. 1895. The species of Dineutes of America north of Mexico. Transactions of the American Entomological Society 22: 279–288.
        Say T. 1825. Descriptions of new species of coleopterous insects inhabiting the United States. Journal of the Academy of Natural Sciences of Philadelphia 5(1): 160-240.
        Sharp D. 1914. The British species of Gyrinus. Entomology Monthly Magazine 50: 128-138.
        Stephens JF. 1835. Illustrations of British entomology; or a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata 5 (App.): 366- 440.
        Thomson CG. 1860. Skandinaviens Coleoptera, synoptiskt bearbetade. Volume II. Lundbergska Boktryckeriet, Lund. 304 pp.
        Wallis JB. 1926. The status of G. piceolus Blatchley (Coleoptera). The Canadian Entomologist 56: 50.
        Zaitsev FA. 1907. Aquatic beetle (Haliplidae, Dytiscidae et Gyrinidae) of Petersburg Province. [In Russian]. Annuaire du Mussee Zologique de L'Academie Imperiale des Sciences de St. Petersbourg 11(1906): 1-46.

    HALIPLIDAE Aubé, 1836*: 14 (Crawling Water Beetles)

    Photos of adult and larva of Brychius hungerfordi (Copyright © 2006 Ethan Bright)

    These are small (<5 mm length) water beetles characterized in the adult form by expanded coxal plates that function as an extension to store air under the elytra (plastron respiration). Although adults have modified tarsi and tibiae for swimming, they are characterized as cumbersome swimmers and are often found clambering on vegetation. Though mostly algivores, the diet of both larva and adult stages is facultatively omnivorous. Eggs are laid on algal food sources, and the three larval stages occur in water. Pupation occurs adjacent to aquatic habitats, with adults emerging to disperse into aquatic habitats. Useful summaries of haliplid biology are given by Matheson (1912), Hickman (1929, 1930) and Roughley (2000).

    Of the approximately 220 described species in all major zoogeographic regions, 20 species in 3 genera have been documented from Michigan, with another 7 occurring nearby based on regional records. Species of Haliplus occur most frequently in ponds and lake margins, and sometimes along slow-moving stream margins, and those of Peltodytes more frequently along stream edges and vegetated banks. The habitat of the endangered species Brychius hungerfordi, known only from a handful of scattered populations in Michigan and Ontario, is restricted to shallow erosional zones of coldwater streams (Strand and Spangler 1994, Mousseau and Rougley 2007). A species identified as Haliplus ruficollis (DeGeer, 1774: 404) from Michigan in the UMMZ collection is probably a misidentification, as this is a Palaearctic species.

      Brychius Thompson 1859*: 11
        Brychius hungerfordi Spangler, 1954*: 114 - Endangered Species (collection in the USA illegal without a U.S. federal permit) Spangler 1954; Strand and Spangler 1994; Legge 1997; UMMZ

      Haliplus Latreille 1802*: 77
        Haliplius (Liaphlus) apostolicus Wallis, 1933*: 46 - MN, WI
        Haliplus (Haliplus) blanchardi Roberts, 1913*: 108 - UMMZ
        Haliplus (Paraliaphlus) borealis LeConte, 1850*: 212 - NMNH (web page database)
        Haliplus (Liaphlus) canadensis Wallis, 1933*: 51 - UMMZ
        Haliplus (Liaphlus) connexus Matheson, 1912: 164 - UMMZ
        Haliplus (Liaphlus) cribrarius LeConte, 1850*: 212 - Hickman 1929: 414-418; Downie and Arnett 1996
        Haliplus (Paraliaphlus) fasciatus Aubé, 1838*: 30 - Downie and Arnett 1996
        Haliplus (Liaphlus) fulvus (Fabricius, 1801*: 271 as Dytiscus) - UMMZ, record at Albertson College of Idaho Collection (as H. subguttatus), ON, BC-MA/NS, widespread north (Roughley 1991)
          (syn.) Dytiscus interpunctatus Marsham, 1802*: 429
          (syn.) Haliplus fulvus var. carlittensis Régimbart, 1901: 323
          (syn.) Haliplus lapponum Thomson, 1856*: 184
          (syn.) Haliplus lapponum var. niger Seidlitz, 1887*: 31
          (syn.) Haliplus pyraeneus Delarouzée, 1857*: 95
          (syn.) Haliplus subguttatus Roberts, 1913*: 100
          (syn.) Haliplus salinarius Wallis, 1933*: 556
          (syn.) Haliplus sparreschneideri Munster, 1922: 15
          (syn.) Haliplus unicolor Fiori, 1914*: 188
          (syn.) Haliplus fulvus var. unicolor Munster, 1922:14
        Haliplus (Haliplus) immaculicollis Harris, 1828*: 164 - Hickman 1929: 409-414; UMMZ; Merritt et al. 2002: 155
          (syn.) Haliplus americanus Aubé, 1838*: 21
          (syn.) Haliplus pallidus Roberts, 1913*: 109
          (syn.) Haliplus robertsi Zimmermann, 1924*: 73
        Haliplus (Paraliaphlus) leopardus Roberts, 1913*: 98 - IL, IN, OH, WI
        Haliplus (Haliplus) longulus LeConte, 1850*: 211 - UMMZ
          (syn.) Haliplus hoppingi Wallis, 1933*: 13
        Haliplus (Liaphlus) nitens LeConte, 1850*: 212 - Vondel 2015
        Haliplus (Paraliaphlus) ohioensis Wallis, 1933*: 27 - IL, IN, OH
        Haliplus (Paraliaphlus) pantherinus Aubé, 1838*: 29 - Downie and Arnett 1996
        Haliplus (Paraliaphlus) tortilipenis Brigham and Sanderson, 1972*: 18 - IL (Cook Co.), MN, WI
        Haliplus (Paraliaphlus) triopsis Say, 1823*: 106 - Hickman 1929: 418-421; UMMZ
        Haliplus (Paraliaphlus) variomaculatus Brigham and Sanderson, 1973*: 157 - IL (Vermillion Co.), may be found in sw. LP

      Peltodytes Regimbart 1879*: 450
        Peltodytes (Neopeltodytes) edentulus (LeConte, 1863*: 21 as Cnemidotus) - Hickman 1929: 408-409; UMMZ, NMNH (web page database)
        Peltodytes (Neopeltodytes) duodecimpunctatus (Say, 1823*: 106 as Haliplus) - Vondel 2015
        Peltodytes (Neopeltodytes) lengi Roberts, 1913*: 119 - Hickman 1929: 400-407; Downie and Arnett 1996
        Peltodytes (Neopeltodytes) muticus (LeConte, 1863*: 21 as Cnemidotus) - UMMZ
        Peltodytes (Neopeltodytes) pedunculatus (Blatchley, 1910*: 204 as Cnemidotus)- Downie and Arnett 1996 - NMNH (web page database)
        Peltodytes (Neopeltodytes) sexmaculatus Roberts, 1913*: 117 - Hickman 1929: 407-408; Downie and Arnett 1996
        Peltodytes (Neopeltodytes) tortulosus Roberts, 1913*: 118 - Downie and Arnett 1996
          (syn.) Peltodytes obesus Peschet, 1923

      References - Taxonomic and Distribution

        Aubé C. 1836-1838. Hydrocanthares, in Dejean PF. (ed.), Iconographie et histoire naturelle des coléoptères d’Europe. Volume V. Méquignon-Marvis, Paris. xi + 416 pp + 46 pls. [Pp. 1-64 (1836), 65-224 (1837), 225-416 (1838).]
        Blatchley WS. 1910. An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of the Rhynchophoral known to occur in Indiana with bibliography and descriptions of new species. Bulletin no. 1 Indiana Department of Geology and Natural Resources. Indianapolis, Indiana, 1386 pp., illus. Reprinted by Nature Publishing Co. in 1910.
        Brigham WU, Sanderson MW 1972. A new species of Haliplus from Illinois and South Dakota. Transactions of the Illinois Academy of Science 65: 17-22. (PDF)
        Brigham WU, Sanderson MW. 1973. Haliplus variomaculatus, a new species from east-central Illinois (Coleoptera: Haliplidae). The Coleopterists Bulletin 27(4): 157-163.
        DeGeer Memoires IV
        Delarouzée C. 1857. Bulletin entomologique 3e Trimestre 1857. Annales de la Société Entomologique de France 5: xciv-xcv.
        Downie NM, Arnett RH. 1996. The beetles of Northeastern North America. Volume I. Introduction; Suborders Archostemata, Adephaga, and Polyphaga, thru Superfamily Cantharoidea. The Sandhill Crane Press: Gainesville, Florida. xiv + 880 pp.
        Fabricius JC. 1801. Systema Eleutheratorum secundum ordines, genera, species; adiectis synonymis, locis, observationibus, descriptionibus. Vol. 1. Bibliopolii Academici Novi, Kiliae. xxiv + 506 pp.
        Fiori A. 1914. Appunti sulla fauna coleotterologica dell’Italia meridionale e della Sicilia, Adephaga. Rivista Coleotterologica Italiana 12:166-191.
        Harris TW. 1828. Contributions to entomology. No. VI. – New England Farmer 7: 164.
        Hickman JR. 1929(1930). Life-histories of Michigan Haliplidae (Coleoptera). Michigan Academy of Science, Arts and Letters 11: 399-424, 9 pls.
        Hickman JR. 1931. Contribution to the biology of the Haliplidae (Coleoptera). Annals of the Entomological Society of America 24: 129-142.
        Latreille PA. 1802. Histoire naturelle, générale et particulière des Crustacés et des Insectes. Familles naturelles et genres. Volume 3. F. Dufart, Paris. xii + 468 pp. (VIEW)
        LeConte JL. 1850. General remarks upon the Coleoptera of Lake Superior, pp. 201-242 + pl. 8 in Agassiz JLR, Cabot JE (eds.), Lake Superior: Its physical character, vegetation and animals. Gould, Kendall & Lincoln, Boston. x + 428 pp. + 8 pls. (PDF)
        LeConte J L. 1863. New species of North American Coleoptera, Part 1. Smithsonian Miscellaneous Collections 6 (167) : 1-86. (PDF)
        Legge J. 1997. 1996 Michigan surveys for the Hungerford's crawling water beetle. Newsletter of the Michigan Entomological Society 42(1): 1.
        Marsham T. 1802. Entomologia Britannica : sistens insecta Britanniae indigena, secundum methodum Linnaenam disposita. Tomus I. Coleoptera. Londini: White, xxxi + 548 p. (PDF)
        Matheson R. 1912. The Haliplidae of North America, north of Mexico. Journal of the New York Entomological Society 20: 156-192, 7 pls. (PDF)
        Mousseau T, Roughley RE. 2007. Taxonomy, classification, reconstructed phylogeny and biogeography of Nearctic species of Brychius Thomson (Coleoptera: Haliplidae). The Coleopterists Bulletin 61(3): 351-397.
        Régimbart M. 1878. Étude sur la classification des Dytiscidae. Annales de la Société Entomologique de France 8: 447–451. (VIEW)
        Roberts CH. 1913. Critical notes on the species of Haliplidae of America north of Mexico with descriptions of new species. Journal of the New York Entomological Society 21(2): 91-123. (PDF)
        Roughley RE. 1991. Family Haliplidae, crawling water beetles. pp. 60–61 in Bousquet Y (ed.), Checklist of Beetles of Canada and Alaska. Agriculture Canada Publication, 1861/E. Ottawa, Ontario. vi + 430 pp.
        Roughley RE. 2000. Haliplidae Aubé, 1836, pp. 138-143 in Arnett RH, Jr., Thomas MC (eds.), American Beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida. 464 pp.
        Say T. 1823. Descriptions of insects of the families of Carabici and Hydrocanthari of Latreille, inhabiting North America. Transactions of the American Philosophical Society, New Series 2(1825)(1): 1-109. (PDF)
        Seidlitz G. 1887. Bestimmungs-Tabelle der Dytiscidae und Gyrinidae des europäischen Faunengebietes. Verhandlungen des Naturforschenden Vereines in Brünn 25(1886): 3-136. (PDF)
        Spanger PJ. 1954. A new species of water beetle from Michigan (Coleoptera: Haliplidae). Entomological News 65:113-117. (PDF)
        Strand RM, Spanger PJ. 1994. The natural history, distribution, and larval description of Brychius hungerfordi Spangler (Coleoptera: Haliplidae). Proceedings of the Entomological Society of Washington 96(2): 208-213.
        Thomson CG. 1856. Öfversigt af de arter inom familjen Dytisci, som blifvit anträffade pa Skandinaviska halfön. Kongliga Svenska Vetenskaps-Akademiens Handlingar 12(1854): 181-237.
        Thomson CG. 1859. Skandinaviens Coleoptera, synoptiskt bearbetade. Tome I. Berlingska Boktryckeriet, Lund. [5] + 290 pp.
        Vondel B, van. 2015. June 22, 2015, World checklist of freshwater Coleoptera: Haliplidae species. World Wide Web electronic publication. Available online at http://fada.biodiversity.be/group/show/64 [accessed 20-July-2017]
        Wallis JB. 1933.
        Revision of the North American species (North of Mexico), of the genus Haliplus, Latreille. Transactions of the Royal Canadian Institute 19: 1-76.
        Zimmermann A. 1924. Die Halipliden der Welt. Entomologische Blätter (für Biologie und Systematik der Käfer) 20 (1): 1-16; (2): 65-80; (3): 129-144; (4): 193-213.

    NOTERIDAE Thomson, 1860*: 34 (Burrowing Water Beetles)

    Photo of Hydrocanthus iricolor (Copyright © 2012 Tom Murray, used with permission)
    Copyright © 2012 Tom Murray
      Hydrocanthus Say, 1823*: 105
        Hydrocanthus iricolor Say, 1823*: 105 - UMMZ; Leng 1920*: 76; Young 1985*: 97; Downie and Arnett 1996*: 268
          (syn.) Hydrocanthus similator Zimmermann, 1928*: 165

      Pronoterus Sharp, 1882*: 263
        Pronoterus semipunctatus (LeConte, 1878*: 595 as Suphis) - LeConte 1878*: 595, Monroe; Young 1954*: 126; Downie and Arnett 1996*: 267

      Suphisellus Crotch, 1873*: 397
        Suphisellus bicolor (Say, 1831*: 33 as Noterus) - IL, IN, OH
          (nom.nud.) Hydroporus gibbulus Dejean, 1833*: 58
        Suphisellus puncticollis Crotch, 1873*: 397 - Young 1954*: 131; Young 1979*:418; Downie and Arnett 1996*: 267

      References
        Crotch GR. 1873. Revision of the Dytiscidae of the United States. Transactions of the American Entomological Society 4: 383-424.
        Dejean PFMA. 1833. Catalogue des coléoptères de la collection de M. le comte Dejean. Paris, 176 pp.
        Downie NM, Arnett RH, Jr. 1996. The beetles of northeastern North America. Vol. 1. The Sandhill Crane Press, Gainesville, Florida. 880 pp.
        LeConte JL. 1878. Descriptions of new species, in Hubbard GH, Schwarz EA, The Coleoptera of Michigan. Proceedings of the American Philosophical Society 17: 593-626.
        Leng CW. 1920. Catalogue of the Coleoptera of America north of Mexico. J.D. Sherman, Mount Vernon, N.Y. 470 pp.
        Say T. 1823. Descriptions of insects of the families of Carabici and Hydrocanthari of Latreille, inhabiting North America. Transactions of the American Philosophical Society, New Series 2(1825)(1): 1-109. (PDF)
        Say T. 1830-1834. Description of new North American insects, and observations on some already described. New Harmony, Indiana: 81 pp (Reprinted in Transactions of the American Philosophical Society 4(1834): 409-470).
        Sharp D. 1878. List of aquatic Coleoptera collected by M. Camille van Volxem in Portugal & Marocco. Annales de la Société Entomologique de Belgique 20(1877): 112-115.
        Thomson CG. 1860. Skandinaviens Coleoptera, synoptiskt bearbetade. Volume II. Lundbergska Boktryckeriet, Lund. 304 pp.
        Young FN. 1954. The water beetles of Florida. University of Florida Studies, Biological Science Series 5: 1-238.
        Young FN. 1979. Water beetles of the genus Suphisellus Crotch in the Americas north of Colombia (Coleoptera: Noteridae). The Southwestern Naturalist 24(3): 409-429.
        Young FN. 1985. A key to the American species of Hydrocanthus Say, with descriptions of new taxa (Coleoptera: Noteridae). Proceedings of the Academy of Natural Sciences of Philadelphia 137(1): 90-98.
        Zimmermann A. 1928. Neuer Beitrag zur Kenntnis der Schwimmkäfer. Wiener Entomologische Zeitung 44: 165-187.
Polyphaga

    CHRYSOMELIDAE Latreille, 1802 (Leaf Beetles)

    The great majority of species in this very large beetle family are terrestrial, but a few taxa, which have specialized on phytophagy upon Nymphaeaceae and other aquatic plants, have become associated with, or adapted to, aquatic habitats (Jolivet and Hawkeswood 1995). The ecology of aquatic and semiaquatic chrysomelids has been summarized by Jolivet and Verma (2002). Larvae of all aquatic species (Donaciinae) use a pair of specialized hooks at the end of the abdomen to tap oxygen from air-containing chambers in plant tissue. Mandibles of the larvae are adapted to macerate plant tissue, and the maxilliary galea and lacinia are adapted to use pharyngeal suction of plant fluids. Larvae construct pupal chambers on their host plants, which is connected to the plant's air cavities. As in the aquatic-adapted Curculionidae, semiaquatic taxa utilize cuticular waxes and hairs as a respiratory plastron or to aid in swimming.Species citations are based on Downie and Arnett (1996), and this list must be considered provisional as no other literature has been consulted, or musuem specimens examined.

      Disonycha Chevrolat, 1844 (Subfamily Galerucinae, Tribe Alticini)
        [not all species in genus are semiaquatic, and I have tried to omit terrestrial species; on Amaranthaceae]
        Disonycha admirabilis Blatchley, 1924 - IN, OH (Downie and Arnett 1996: 1370)
        Disonycha alternata (Illiger), 1807 Haltica (orig.) - Downie and Arnett 1996: 1370 (on Salix, willow; perhaps periaquatic)
        Disonycha latifrons laticollis Schaeffer, 1919 - Downie and Arnett 1996: 1370 (found in wetlands on goldenrod; perhaps periaquatic)
        Disonycha pensylvanica (Illiger, 1807) Haltica (orig.) - (on Polygonum spp.)
        Disonycha procera Casey, 1884 - Downie and Arnett 1996: 1370 (on Polygonum spp.)
        Disonycha uniguttata (Say, 1824) Altica (orig.) - Downie and Arnett 1996: 1370

      Donaciella Rietter, 1920 (Subfamily Donaciinae)
        Donaciella pubicollis (Suffrian, 1872) Donacia (orig.) - Downie and Arnett 1996: 1289 (on Phragmites spp., reed-grass, and Nymphaea spp., water lilies)
          (orig.) Donacia pubicollis Suffrain, 1872:21
          (syn.) Donacia pubicollis Crotch (1872:21, not Suffrian, 1872:21)

      Donacia Fabricius, 1775 (Subfamily Donaciinae)
        Donacia (Donaciomima) biimpressa Melsheimer, 1847 - Downie and Arnett 1996:1298 (on Carex spp., Scirpus spp., bulrushes)
          (syn.) Donacia aurichalcea Melsheimer, 1847:159)
          (syn.) Donacia torosa LeConte, 1851:313
        Donacia (Donaciomima) caerulea Olivier, 1795 - Downie and Arnett 1996:1296 (on Sagittaria spp.; also on Acorus calamus sweet flag, and Pontederia cordata, pickerel weed); Pine Creek, 10 mi S Battle Creek, MI (Kölsch & Pedersen 2008)
          (syn.) Donacia aequalis Say, 1824:248
          (syn.) Donacia confusa, Lacordaire 1845:109, not LeConte, 1851
          (syn.) Donacia confusa LeConte 1851:313, not Lacordaire, 1845
        Donacia (Donaciomima) cazieri Marx, 1957 - Munising Tourist Park (Kölsch & Pedersen 2008) (Downie and Arnet 1996:1298, on Carox limosa, mud sedge)
        Donacia (Donacia) cincticornis Newman, 1838 - Downie and Arnett 1996: 1295 (on Nymphaea spp. and Nuphar spp., water lilies, Potamogeton spp. pondweed, Nymphoides spp., floating heart)
          (syn.) Donacia lucida Lacordaire, 1845:106
          (syn.) Donacia rufipennis Lacordaire, 1845:110
          (syn.) Donacia pulchella LeConte, 1851:312
          (syn.) Donacia antillarum Suffrain, 1864:282
          (syn.) Donacia cincticornis var. tryphera Schaeffer, 1925:81
          (syn.) Donacia cincticornis var. tenuis Schaeffer, 1925:82
        Donacia (Donaciomima) confluenta Say, 1827 - Downie and Arnett 1996:1297 (on Sparganium spp., bur-reeds); Pine Creek, 10 mi S Battle Creek, MI; ditch along Mitchell Lake Rd., MI (Kölsch & Pedersen 2008)
        Donacia (Donaciomima) cuprea Kirby, 1837 - Downie and Arnett 1996:1296 (on Scirpus occidentalis)
          (syn.) Donacia curticollis Knab, 1905:122
        Donacia (Donaciomima) distincta Leconte, 1851 - Downie and Arnett 1996:1298 (on Carex spp. sedges, Scirpus spp., bulrushes)
        Donacia (Donaciomima) fulgens Leconte, 1851- Downie and Arnett 1996:1296-1297 (on Sparganium angustifolium, narrow leafed bur-reed, Pontederia cordata, pickerel weed); small lake near Lake City, MI (Kölsch & Pedersen 2008)
        Donacia (Donaciomima) hirticollis Kirby, 1837 - Downie and Arnett 1996:1296 (on Potamogeton spp., other aquatic plants); small lake near Lake City, MI (Kölsch & Pedersen 2008)
          (syn.) Donacia rudicollis Lacordaire, 1845:108
        Donacia (Donaciomima) liebecki Schaeffer, 1919 - Downie and Arnett 1996:1297 (on Nymphaea spp., Nymphar spp., Orontium aquaticum, floating arum)
          (syn.) Donacia pallipes Lacordaire, 1845:149
        Donacia (Donaciomima) limonia Schaeffer, 1925 - IN, ON, may occur in s. LP (Downie and Arnett 1996:1298; on Carex spp., sedges, and Scirpus spp., bulrushes)
        Donacia (Donaciomima) magnifica Leconte, 1851 - ON, MN, widespread (Downie and Arnett 1996:1297 (on Potamogeton spp., pond weeds, Sagittaria spp., arrow leaf)
        Donacia (Donaciomima) megacornis Blatchley, 1910 - Downie and Arnett 1996:1297 (on Nymphaea spp., Nymphar spp.)
          (syn.) Donacia megatocera Weise 1913:118
        Donacia (Donacia) palmata Olivier, 1795 - Downie and Arnett 1996: 1294 (on Nymphaea spp. and Nuphar spp., water lilies); ; small lake near Lake City, MI (Kölsch & Pedersen 2008)
          (syn.) Donacia claudicans Germar, 1821:173
          (syn.) Donacia palmata angustipes Marx, 1957:237
        Donacia (Donacia) parvidens Schaeffer, 1919 - Downie and Arnett 1996: 1295 (on Nymphaea spp. and Nuphar spp., water lilies)
        Donacia (Donacia) piscatrix Lacordaire, 1845 - Downie and Arnett 1996: 1294-1295 (on Nymphaea spp. and Nuphar spp., water lilies); shallow lake E Tacoma Lake, MI (Kölsch & Pedersen 2008)
          (syn.) Donacia carolina Lacordaire, 1845:114
          (syn.) Donacia cuprea Melsheimer, 1847:158, not Kirby, 1837
          (syn.) Donacia alutacea LeConte, 1851:311
          (syn.) Donacia congener LeConte, 1851:310
        Donacia (Donaciomima) porosicollis Lacordaire, 1845 - MN, ON, expect in Isle Royale, w UP (Downie and Arnett 1996: 1298; on Carex spp. sedges, and Scirpus spp. nut rushes)
        Donacia (Donacia) proxima Kirby, 1837 - Downie and Arnett 1996: 1295 (on Nymphaea spp. and Nuphar spp., water lilies)
          (syn.) Donacia espiscopalis Lacordaire, 1845:105
          (syn.) Donacia california LeConte, 1861:357
          (syn.) Donacia caerulea, sensu Schaeffer 1925 and Marx 1957
        Donacia (Donaciomima) pubescens Leconte, 1868 - Downie and Arnett 1996:1296 (on Scirpus occidentalis)
        Donacia (Donacia) rufescens Lacordaire, 1845 - IN, OH, ON (Downie and Arnett 1996: 1295 (on Nymphaea spp. and Nuphar spp., water lilies), widespread east)
        Donacia (Donaciomima) rugosa Leconte, 1851 - Downie and Arnett 1996:1298 (on Pontederia cordata, pickerel weed)
          (syn.) Donacia aenea Ahrens, 1810:21 (not Hoppe 1875:44, not Gistel 1857:524)
          (syn.) Donacia quadricollis Say, 1827:282
          (syn.) Donacia aerea Lacordaire, 1845:148 (not Schrank, 1798)
          (syn.) Donacia subtilis magistrigata Mead, 1938:113
        Donacia (Donaciomima) subtilis Kunze, 1818 - Downie and Arnett 1996:1297 (on Sparganium spp., bur-reeds); Kalamazoo River, 4 mi E Douglas, MI; Fawn River, 1 mi E Constantine, MI (Kölsch & Pedersen 2008)
        Donacia (Donacia) texana Crotch, 1873 -Downie and Arnett 1996: 1295-1296 (on Nymphaea spp. water lilies)
          (syn.) Donacia texana var. minor Schaeffer, 1925:72
        Donacia (Donaciomima) tuberculifrons Schaeffer, 1919 - Downie and Arnett 1996:1296 (on Scirpus paludosus, Sparganium spp., Carex spp., Acorus calamus sweet flag)

      Hydrothassa Thomson, 1866
        [Ranunculus and marsh vegetation]
        Hydrothassa boreella Schaeffer, 1928 - widespread, expect northern Michigan (Downie and Arnett 1996: 1346)
        Hydrothassa obliquata (Leconte, 1866) Prasocuris (orig.) - IN, OH (Downie and Arnett 1996: 1346)
        Hydrothassa ovalis (Blatchley, 1910) Prasocuris (orig.) - Downie and Arnett 1996: 1346, on Carex spp.
        Hydrothassa vittata (Olivier, 1808) Helodes (orig.) - IN, OH, widespread (Downie and Arnett 1996: 1346)
          (syn.) Hydrothassa trivittata (Say)
          (syn.) Hydrothassa varipes (LeConte)

      Neohaemonia Székessy, 1941
        Neohaemonia flagellata Askevold, 1988 - IL, IN, ON, MN (Downie and Arnett 1996: 1289; widespread, on Potamogeton spp.)
        Neohaemonia melsheimeri (Lacordaire, 1845) Haemonia (orig.) - Downie and Arnett 1996: 1289; on Potamogeton spp.
        Neohaemonia minnesotensis Askevold, 1988 - Downie and Arnett 1996: 1289; on Potamogeton spp.
        Neohaemonia nigricornis (Kirby, 1837) Haemonia (orig.) - MN, ON, WI, widespread (Downie and Arnett 1996: 1288-1289; on Potamogeton spp.)
          (orig.) Haemonia nigricornis Kirby, 1837:222)
          (syn.) Haemonia americana Guérin-Méneville, 1844:249

      Plateumaris Thomson, 1859
        Plateumaris aurifera (LeConte, 1851) Donacia - MN (Askevold 1991: 155; expect in northern UP, on Carex, Scirpus and Eleocharis)
        Plateumaris diversa (Schaeffer, 1925) Donacia - Askevold 1991: 40, 151 (ESGR, Livinston Co.; on sedges in moist meadows)
          (syn.) Donacia harrisi Blatchley, 1910:1105 (misid.)
        Plateumaris flavipes (Kirby, 1837) - Donacia - Askevold 1991: 156 (on Carex, Scirpus and Eleocharis)
          (syn.) Donacia wallisi Schaeffer, 1925
        Plateumaris frosti (Schaeffer, 1925) - Askevold 1991: 164 (on Carex esp. stricta Lam., Scirpus and Eleocharis), Kölsch & Pedersen 2008: 938 (Soldier Lake [46.349N, -84.867W], Chippewa Co.)
        Plateumaris fulvipes (Lacordaire, 1845) - Askevold 1991: 158 (on Carex, Scirpus and Eleocharis)
          (syn.) Donacia fulvipes Dejean, 1837 (nom.nudum)
        Plateumaris germari (Mannerheim, 1843) - Donacia - Askevold 1991: 157 (on Carex, Scirpus, Juncus and Eleocharis)
          (syn.) Donacia flavipennis Mannerheim, 1843
          (syn.) Donacia dives LeConte, 1851
          (syn.) Donacia serricauda Schaeffer, 1919
          (syn.) Donacia femoralis LeConte, 1851 (misid.)
        Plateumaris metallica (Ahrens, 1810) - Donacia - Askevold 1991: 160 (on Carex, Scirpus, and Juncus affinis R. Brown, adventitious on other plants)
          (syn.) Donacia femoralis, Kirby (1837: 223), Lacordaire, 1845
          (syn.) Donacia cataractae Newman (1838: 391)
          (syn.) Donacia parva Lacordaire (1845: 190)
          (syn.) Donacia indica Melsheimer (1847: 158)
          (syn.) Donacia nana Melsheimer (1847: 160)
          (syn.) Donacia gentilis LeConte (1851: 314)
        Plateumaris nitida (Germar, 1811) - Donacia - Askevold 1991: 163 (on Carex, Scirpus and Eleocharis, adventitious on other plants), Kölsch & Pedersen 2008: 938 (M-115 at Middle Branch River, [44.088N, -85.231W], Osceola Co.)
          (syn.) Donacia emarginata Kirby, 1837, var. pacifica Schaeffer, 1925
          (syn.) Donacia juncina Couper, 1864
        Plateumaris pusilla (Say, 1827) - Donacia - Askevold 1991: 153 (on Carex, Scirpus, Juncus and Eleocharis, adventitious on other plants)
          (syn.) Donacia rugifrons Newman, 1838 (nom.nudum), Lacordaire, 1845
          (syn.) Donacia pyritosa LeConte, 1857
          (syn.) Donacia cuprea Crotch, 1873:31 (misid.)
        Plateumaris robusta (Schaffer, 1919) - Donacia - Askevold 1991: 154 (on Carex, Scirpus and Eleocharis)
        Plateumaris rufa (Say, 1827) - Donacia - Askevold 1991: 152 (on Carex and Scirpus)
          (syn.) Donacia sulcicollis Dejean, 1837 (nom.nudum), Lacordaire, 1845
          (syn.) Donacia affis Kirby, 1837
          (syn.) Donacia chalcea Lacordaire, 1845
          (syn.) Donacia kirbyi Lacordaire, 1845
          (syn.) Donacia jacunda LaConte, 1850 (nom.nudum), 1851
        Plateumaris shoemakeri (Schaeffer, 1925) - Donacia - Askevold 1991: 159 (on Acorus calamus L., accidental or adventitious on other plants)
          (syn.) Donacia flavipes var. shoemakeri Schaeffer, 1925
          (syn.) Donacia flavipes var. lodingi Schaeffer, 1925
          (misid.) Plateumaris chalcea, Askevold, 1990
        Plateumaris schaefferi Askevold, 1991 - Askevold 1991: 162

      Prasocuris Latreille, 1802
        Prasocuris phellandrii (Linnaeus, 1758) Chrysomela (orig.) - IN, OH, ON, widespread (Downie and Arnett 1996: 1345, on Caltha spp. (marsh marigold) and Sium suave (water parsnip)

      Pyrrhalta Joannis, 1866
        [not aquatic]
        Pyrrhalta (Tricholochomaea) alni (Fall, 1924) Galerucella (orig.) - Downie and Arnett 1996:1352 (on Alnis, alder; periaquatic)
        Pyrrhalta (Tricholochomaea) decora (Say, 1824) Galleruca (orig.) - Downie and Arnett 1996 (on Salix, willow; perhaps periaquatic)
        Pyrrhalta (Neogalerucalla) quebecensis (Brown, 1938) Galerucella (orig.) - Downie and Arnett 1996:1350 (on Potentialla palustris, cinquefoil; periaquatic)
        Pyrrhalta (Tricholochomaea) ribicola (Brown, 1938) Galerucella (orig.) - Downie and Arnett 1996:1352 (on Ribes, gooseberry; periaquatic, especially bogs)
        Pyrrhalta (Tricholochomaea) tuberculata (Say, 1824) Galleruca (orig.) - Downie and Arnett 1996:1352, (on Salix, willow; perhaps periaquatic)

    CURCULIONIDAE Latreille, 1802 (True Weevils)

    Photo of Auleutes nebulosus by Tom Murray (Copyright © 2011, used with permission).
    Copyright © 2011 Graham Montgomery, used with permission

    This list is based on Downie and Arnett (1996) and must be considered provisional. No museum specimens or other literature citations have been consulted.

      Auleutes Dietz, 1896, 1802 (Subfamily Ceutorhynchinae)
        (not all species listed herein are aquatic/semiaquatic)
        Auleutes asper (LeConte, 1876) Coeliodes (orig.) - IN
        Auleutes atar (LeConte, 1876) Coeliodes (orig.) - IN
        Auleutes epilobii (Paykull, 1800) Curculio (orig.) - Downie and Arnett 1996
        Auleutes marionis Fall, 1913 - IN
        Auleutes nebulosus (LeConte, 1876) Coeliodes - Downie and Arnett 1996
        Auleutes subfasciatus Dietz, 1896 - Downie and Arnett 1996
        Auleutes tenuipes (LeConte), 1876 Coeliodes - IN, ON

      Bagous Germar, 1817 (Subfamily Bagoinae)
        Bagous americanus LeConte, 1876 - Tanner 1943:19-20 (also B. longirostrus? 18-19); Downie and Arnett 1996:1535
        Bagous atratus Blatchley, 1916 - Tanner 1943:23-24; Downie and Arnett 1996:1535
        Bagous bituberosa LeConte, 1876 - IN (widespread, Tanner 1943:29-30; Downie and Arnett 1996:1536)
        Bagous blanchardi Blatchley, 1916 - Downie and Arnett 1996:1535
        Bagous calvifrons LeConte, 1876 - Tanner 1943:21-22; Downie and Arnett 1996:1535
        Bagous magister LeConte, 1876 - Tanner 1943:22; Downie and Arnett 1996:1535
        Bagous mamillatus Say, 1831 - Tanner 1943:16-18; Downie and Arnett 1996:1534-1535
        Bagous nebulosus LeConte, 1876 - Tanner 1943:25; Downie and Arnett 1996:1535
        Bagous obliquus LeConte, 1876 - Downie and Arnett 1996:1534
        Bagous planatus LeConte, 1876 - Tanner 1943:12; Downie and Arnett 1996:1534
        Bagous puritanus Blatchley, 1916 - IN, IL (Downie and Arnett 1996:1535-1536)
        Bagous pusillus LeConte, 1876 - IL, IN (Downie and Arnett 1996:1536)
        Bagous restrictus LeConte, 1876 - IL, IN, MN, OH, WI (Tanner 1943:28; Downie and Arnett 1996:1535)
        Bagous tanneri O'Brien, 1979 - Downie and Arnett 1996:1535
        Bagous transversus LeConte, 1876 - Tanner 1943:31-32; Downie and Arnett 1996:1535
          (syn.) Bagous carinatus Blatchley, 1925

      Brachybamus Germar, 1836 (Subfamily Erirhininae)
        Brachybamus electus Germar, 1836 - IN (widespread, on Sagittaria; Downie and Arnett 1996:1538)

      Euhrychiopsis Dietz, 1896 (Subfamily Ceutorhynchinae)
        Euhrychiopsis lecontei (Dietz, 1896) Phytobius (orig.) - Creed 1998

      Lissorhopterus LeConte 1876 (Subfamily Erirhininae)
        Lissorhopterus buchanani Kuschel, 1952 - Downie and Arnett 1996:1537
        Lissorhopterus lacustris Kuschel, 1952 - Downie and Arnett 1996:1537
        Lissorhopterus oryzophilus Kuschel, 1952 - Downie and Arnett 1996:1537 (rice water weevil)

      Listronotus Jekel, 1865 (Subfamily Cylydrorhininae)
        [some species listed herein may not be aquatic or semi-/peri-aquatic]
        Listronotus appendiculatus (Boheman, 1842) Listroderes (orig.) - Downie and Arnett 1996:1503
        Listronotus caudatus (Say, 1824) Rhynchaenus (orig.) - Downie and Arnett 1996:1501-1502
        Listronotus delumbis (Gyllenhal, 1834) Listroderes (orig.) - Downie and Arnett 1996:1505
        Listronotus dorsalis (Dietz, 1889) Macrops (orig.) - IN, IL, MN (Downie and Arnett 1996:1504)
        Listronotus echinatus (Dietz, 1889) Macrops (orig.) - Downie and Arnett 1996:1504-1505
        Listronotus echinodori O'Brien, 1977 - MN, OH (Downie and Arnett 1996:1503)
        Listronotus frontalis LeConte, 1876 - IN, IL, MN, OH, ON, WI (Downie and Arnett 1996:1503)
        Listronotus grypidiodes (Dietz, 1889) Macrops (orig.) - IN, MN (Downie and Arnett 1996:1504)
        Listronotus humilus (Gyllenhal, 1834) Listroderes (orig.) - Downie and Arnett 1996:1506
        Listronotus laramiensis (Angell, 1893) Macrops (orig.)
        Listronotus maculicollis (Kirby, 1837) Macrops (orig.) - Downie and Arnett 1996:1505
        Listronotus montanus (Dietz, 1889) Macrops (orig.) - IN, IL, MN
        Listronotus sparsus (Say, 1831) Listroderes (orig.) - Downie and Arnett 1996:1505
        Listronotus squamiger (Say, 1831) Listroderes (orig.)- Downie and Arnett 1996:1502
        Listronotus tuberosus LeConte, 1876 - Downie and Arnett 1996:1502
        Listronotus vitticollis (Kirby, 1837) Macrops (orig.) - ON, south (Downie and Arnett 1996:1504)

      Lixellus LeConte, 1876 (Subfamily Cylydrorhininae)
        Lixellus filiformis LeConte, 1876 - IN, MN (Downie and Arnett 1996:1506)
        Lixellus hubbardi (LeConte, 1876) Anchodemus (orig.) - Downie and Arnett 1996:1506
        Lixellus lutulentus (Boheman, 1843) Erirhinus (orig.) - Downie and Arnett 1996:1506

      Lixus Fabricius, 1801 (Subfamily Cleoninae)
        Lixus caudifer LeConte, 1876 - IL, IN (Downie and Arnett 1996:1513)
        Lixus concavus Say, 1831 - Downie and Arnett 1996:1514
        Lixus musculus Say, 1831 - Downie and Arnett 1996:1514
        Lixus parcus LeConte, 1876 - IL, IN, ON (Downie and Arnett 1996:1514)
        Lixus punctinasus LeConte, 1876 - IL, OH, WI (Downie and Arnett 1996:1514)
        Lixus rubellus Randall, 1838 - Downie and Arnett 1996:1513
        Lixus sobrinus Casey - IL, IN, WI (Downie and Arnett 1996:1515)

      Notiodes Schönherr, 1838 (Subfamily Erirhininae)
        Notiodes ovalis (LeConte, 1876) Endalus (orig.) - IN, OH, ON, MN (Downie and Arnett 1996:1538)
        Notiodes cribricollis (LeConte, 1876) Endalus (orig.) - widespread (Downie and Arnett 1996:1538)
        Notiodes limatulus (Gyllenhal, 1836) Notiophilus (orig.) - Downie and Arnett 1996:1538

      Onychylis LeConte, 1876 (Subfamily Erirhininae)
        Onychylis longulus LeConte, 1876 - Downie and Arnett 1996:1539
        Onychylis nigrirostris (Boheman, 1843) Notiodes (orig.) - Downie and Arnett 1996; 1539

      Phytobius Schönherr, 1833 (Subfamily Ceutorhynchinae)
        Phytobius leucogaster (Marsham, 1802) Curculio (orig.) - Downie and Arnett 1996 (introduced)

      Rhinoncus Schönherr, 1825 (Subfamily Ceutorhynchinae)
        Rhinoncus asperulus (Dietz, 1896) Pelenomus (orig.) - Downie and Arnett 1996
        Rhinoncus fuliginosus (Dietz, 1896) Mecopeltus (orig.) - IN (widespread)
        Rhinoncus gracilipes (Dietz, 1896) Pelenomus (orig.) - widespread
        Rhinoncus squamosus (LeConte, 1876) Pelenomus (orig.) - Downie and Arnett 1996
        Rhinoncus sulcicollis (Fåhraeus, 1843) Phytobius (orig.) - Downie and Arnett 1996

      Stenopelmus Schönherr, 1836 (Subfamily Erirhininae)
        Stenopelmus rufinasus Gyllenhal, 1836 - IN (widespread; Downie and Arnett 1996:1539)

      Tanysphyrus Germar, 1817 (Subfamily Erirhininae) - on Lemna sp. (duckweed)
        Tanysphyrus lemnae (Fabricius) - Downie and Arnett 1996
        Tanysphyrus ater Blatchley - Downie and Arnett 1996

    DRYOPIDAE Erichson, 1857 (Dryopid Beetles)

      Helichus Erichson, 1847
        Helichus basalis LeConte, 1852 - NMNH (web page database); see also Musgrave 1935:140-141
        Helichus fastigiatus (Say, 1824) - UMMZ (Washtenaw Co., Ann Arbor, 20-July-1901, Mus. Cat. 1634, det./ver. FNYoung)
        Helichus lithophilus (Germar, 1824*) - UMMZ (Washtenaw Co., Ann Arbor, 04-Aug-1916, det. H. Nelson 1973; Oakland Co., 04-July-1909, det. H. Nelson), NMNH (web page database); see also Musgrave 1935:140
        Helichus striatus LeConte, 1852 - UMMZ (Cheboygan Co., Cheboygan, 08-Jun-1927, Sherman Moore 1029, det. FNYoung; Livingston Co., ES George Reserve, 855b & 833a coll. IJCantrall & FNYoung), NMNH (web page database); see also Musgrave 1935:142-143

      Pelonomus Erichson, 1847
        Pelonomus obscurus LeConte - IL, IN, ON (widespread central/east)

    ELMIDAE Curtis, 1830*: 294 (Riffle Beetles)

    These are small beetles, with adults of about 2-3 mm in length. All larvae and adults of our species are aquatic and herbivorous, and live on rocks and logs, slowly walking on the surface of their preferred substrate. Adults, after emerging from the pupa on land, engage in a dispersal flight and enter the water. Juveniles occur together with adults in the water, and are easy to collect. Larvae are more mobile than adults, with anal gills protruding an air bubble that provides bouyancy as well as functioning as a source of dissolved oxygen. Similarly, emerging adults, upon re-entering the water, take a thin sheet of air under the plastron into the water and maintain their oxygen from the surrounding water.The larval stage of 5-6 instars usually requires 2-3 years, depending on species, size and temperature. Adults mate in water, and females attach their eggs singly or in small clusters to rock or vegetation. An excellent summary of the biology of this group remains Brown (1987).

    Adult Macronychus glabratus (Copyright © 2005 Ethan Bright)
    Larva of Dubiraphia sp. (Copyright © 2005 Ethan Bright)
      Ancyronyx Erichson, 1847*: 522 (Subfamily Elminae, Tribe Ancyronychini)
        Ancyronyx variegatus (Germar, 1824*: 89 as Macronychus.) - Brown 1983:3; Bright, pers. obs., Rouge River, Wayne Co.
          (syn.) Elmis cincta Say, 1825*: 186
          (syn.) Hydrochus parumoculatus Hardy, 1852: 270

      Dubiraphia Sanderson, 1954*: 4 (Subfamily Elminae, Tribe Elmini)
        Dubiraphia bivittata (LeConte, 1852*: 44 as Elmis) - see also Brown 1983:4
        Dubiraphia minima Hilsenhoff, 1973*: 59 - Yanoviak and McCafferty 1996:205; also IN, OH, WI (Brown 1983:4)
        Dubiraphia quadrinotata (Say, 1825*: 187 as Elmis) - UMMZ (Rice Creek, Calhoun Co., 01-Aug-1949, leg. V.S. Pratt, det. Sanderson, MW 1950), NMNH (web page database); see also Brown 1983:4
        Dubiraphia robusta Hilsenhoff, 1973*: 59 - WI, expect in w. UP (Hilsenhoff 1973:59; Brown 1983:4)
        Dubiraphia vittata (Melsheimer, 1844*: 99 as Elmis) - Brown 1983:5; NMNH (web page database)

      Macronychus Müller, 1806*: 207 (Subfamily Elminae, Tribe Macronychini)
        Macronychus glabratus Say, 1825*: 187 - UMMZ; Bright coll.; Brown 1983:12; Yanoviak and McCafferty 1996:205
          (syn.) Macronychus lateralis Melsheimer, 1844*: 99

      Microcylloepus Hinton, 1935*: 178 (Subfamily Elminae, Tribe Elmini)
        Microcylloepus pusillus (LeConte, 1852*: 44 as Stenelmis) - ON, WI, expect in w. UP (Brown 1983:6)

      Optioservus Sanderson, 1954*: 8 (Subfamily Elminae, Tribe Elmini)
        Optioservus fastiditus (LeConte, 1850*: 218 as Limnius) - Brown 1983:8; Yanoviak and McCafferty 1996:205; UMMZ (Rice Creek, Calhoun Co., 01-Aug-1949, leg. V.S. Pratt, det. Sanderson, MW 1950; Spring Brook, Kalamazoo Co., det. Sanderson, MW 1950)
        Optioservus trivittatus (Brown, 1930a*: 91 as Limnius) - Brown 1983:9; Yanoviak and McCafferty 1996:205; UMMZ (Rice Creek, Calhoun Co., 01-Aug-1949, leg. V.S. Pratt, det. Sanderson, MW 1950)

      Stenelmis Dufour, 1835*: 158 (Subfamily Elminae, Tribe Elmini)
        Stenelmis antennalis Sanderson, 1938*: 695 - MN, WI
        Stenelmis bicarinata LeConte, 1852*: 44
          (syn.) Stenelmis maerkelii Motschulsky, 1854*: 12
          (syn.) Stenelmis vittipennis Zimmerman, 1869*: 259
        Stenelmis crenata (Say, 1824: 181 as Elmis) - Brown 1983:10; Yanoviak and McCafferty 1996:205
          (syn.) Stenelmis sordida Motschulsky, 1859: 51
        Stenelmis decorata Sanderson, 1938*: 701 - Schmude 1992
        Stenelmis douglasensis Sanderson, 1938*: 685 - Brown 1983:10 (holotype from Douglas Lake, Cheboygan Co.)
        Stenelmis fuscata Blatchley, 1925*: 164 - WI
        Stenelmis grossa Sanderson, 1938*: 686 - Schmude 1992
        Stenelmis lignicola Schmude & Brown, in Schmude, Barr & Brown 1992*: 583 - Kalamazoo Co., Gull Lake Biological Station, specimen at Cook Collection - Schumde et al. 1992
          (syn.) Stenelmis convexula Brown, 1956*: 38
          (syn.) Stenelmis markelii [sic] Finni and Skinner 1975*: 390
          (syn.) Stenelmis crenata, S. sulcata Patrick 1961*: 255
          (syn.) Stenelmis bicarinata, S. convexula, S. crenata, S. decorata (in part), S. hungerfordi (in part), S. sulcata Patrick et al. 1967*: 378-379
          (syn.) Stenelmis spp. Benke 1984*: 40 (in part)
        Stenelmis mera Sanderson, 1938a*: 682 - Schmude 1992
        Stenelmis musgravei Sanderson, 1938*: 698 - see also Brown 1983:11
        Stenelmis quadrimaculata Horn, 1870*: 40 - Brown 1983:11
          (syn.) Stenelmis sulcata Blatchley, 1910*: 681
          (syn.) Stenelmis blatchleyi Musgrave, 1933*: 57
        Stenelmis sandersoni Musgrave, 1940*: 48
          (syn.) Stenelmis tarsalis Sanderson, 1938*: 675
        Stenelmis sexlineata Sanderson, 1938*: 663 - Schmude 1992

      References - Elmidae

        Benke AC, Van Arsdall TC, Jr., Gillespie DM, Parrish FK. 1984. Invertebrate productivity in a subtropical blackwater river: the importance of habitat and life history. Ecological Monographs 54: 25-63.
        Blatchley WS. 1910. An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of the Rhynchophoral known to occur in Indiana with bibliography and descriptions of new species. Bulletin no. 1 Indiana Department of Geology and Natural Resources. Indianapolis, Indiana, 1386 pp., illus. Reprinted by Nature Publishing Co. in 1910.
        Blatchley WS. 1925. Notes on the distribution and habits of some Florida Coleoptera with descriptions of new species. The Canadian Entomologist 57: 160·168.
        Brown HP. 1956. Riffle beetles in Oklahoma. Proceedings of the Oklahoma Academy of Science 36: 38.
        Brown HP. 1972. Aquatic dyropoid beetles (Coleoptera) of the United States. Biota of Freshwater Ecosystems Identification Manual No. 6. United States Environmental Protection Agency Project # 18050, Contract # 14-12-894, April 1972. x + 82 pp.
        Brown HP. 1983. A catalog of the Coleoptera of America north of Mexica. Family: Elmidae. United States Department of Agriculture Handbook 529-50. x + 23 pp. (PDF)
        Brown WJ. 1930a. New species of Coleoptera I. The Canadian Entomologist 62: 87-92.
        Curtis J. 1930. British entomology, vol. 7. Unnumbered pages, pls. 290-337. Author, London.
        Dufour L. 1835. Recherches anatomiques et considérations entomologiques sur les insectes coléoptères des genres Macronique et Elmis. Annales des Sciences Naturelles, Series Zoologie 2(3): 151-174, pls. 6-7.
        Erichson WF. 1847. Naturgeschichte der Insecten Deutschlands. Abtheilung 1, Coleoptera., Dritter Band, Vierte Lieferung, pp. 481-640. Nicolaische Buchhandlung, Berlin. (PDF)
        Finni GR, Skinner BA. 1975. The Elmidae and Dryopidae (Coleoptera: Dryopoidea) of Indiana. Journal of the Kansas Entomological Society 48: 388-395.
        Germar EF. 1824. Insectorum species novae aut minus cognitae, descriptionibus illustratae. Volumen primum, Coleoptera. Impensis J.C. Hendelii et Filii, Halae. xxiv + 624 pp. (PDF)
        Hilsenhoff WL. 1973. Notes on Dubiraphia (Coleoptera: Elmidae), with the description of five new species. Annals of the Entomological Society of America 66(1): 55-61.
        Hilsenhoff WL, Schmude KL. 1992. Riffle beetles of Wisconsin (Coleoptera: Dryopidae, Elmidae, Lutrochidae, Psephenidae) with notes on distribution, habitat, and identification. The Great Lakes Entomologist 25(3):191-209.
        Hinton HE. 1935. Notes on the Dryopoidea (Col.). Stylops 4(8): 169-179.
        Horn GH. 1870. Synopsis of the Parnidae of the United States. Transactions of the American Entomological Society 3: 29-42.
        LeConte JL. 1850. General remarks upon the Coleoptera of Lake Superior, pp 201-242 in Agassiz L, LeConte JL, Lake Superior: Its physical character, vegetation, and animals, compared with those of other and similar regions. Gould, Kendall and Lincoln, Boston, 428 pp., illus.
        Le Conte JL. 1852. Synopsis of the Parnidae of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 6: 41-45.
        Melsheimer FE. 1844. Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 2: 98-118.
        Motschulsky TV, von. 1854. Voyages. Lettre de M. de Motschulsky a M. Menetries. Etudes Entomologiques, pt. 3, pp. 1-15.
        Müller PWJ. 1806. Macronychus, Krallenkäfer. Eine neue Käfergattung. Magazin für Insektenkunde 5: 207-220.
        Musgrave PN. 1933. New species of Helmidae (Coleoptera). Proceedings of the Entomological Society of Washington 35(4): 54·57.
        Musgrave PN. 1940. A new name: Elmidae (Coleoptera). Proceedings of the Entomological Society of Washington 42(2): 48.
        Patrick R. 1961. A study of the numbers and kinds of species found in rivers in eastern United States. Proceedings of the Academy of Natural Sciences of Philadelphia 113: 215-258.
        Patrick R, Cairns J, Jr., Roback SS. 1967. An ecosystematic study of the fauna and flora of the Savannah River. Proceedings of the Academy of Natural Sciences of Philadelphia 118: 109-407.
        Sanderson MW. 1938. A monographic revision of the North American species of Stenelmis (Dryopidae: Coleoptera). The University of Kansas Science Bulletin 25(22): 635-717.
        Sanderson MW. 1954. A revision of the Nearctic genera of Elmidae (Coleoptera). Journal of the Kansas Entomological Society 27: 1-13.
        Say T. 1825. Descriptions of new species of coleopterous insects inhabiting the United States. Journal of the Academy of Natural Sciences of Philadelphia 5(1): 160-240.
        Schmude KL. 1992. Revision of the riffle beetle genus Stenelmis (Coleoptera: Elmidae) in North America, with notes on bionomics. Ph.D. dissertation, University of Wisconsin. 388 pp.
        Schmude KL, Barr CB, Brown HP. 1992. Stenelmis lignicola and Stenelmis xylonastis, two new North American species of wood-inhabiting riffle beetles (Coleoptera: Elmidae). Proceedings of the Entomological Society of Washington 94(4): 580-594.
        Zimmermann CCA. 1869. Synonymical notes on Coleoptera of the United States, with descriptions of new species, from the MSS of the late Dr. C. Zimmermann. Edited by John L. LeConte, M.D. Transactions of the American Entomological Society 2: 243- 259.

    HYDRAENIDAE Mulsant, 1844*: 50 (Minute Moss Beetles)
    The information contained on this list is tentative, based on few observations of museum specimens (currently being assessed) and literature citations (Perkins 1980).

      Gymnochthebius d'Orchymont, 1943b*: 38
        Gymnochthebius nitidus (LeConte, 1850*: 217 as Ochthebius) - Perkins 1980: 274-277; NMNH (web page database); holotype specimen from Eagle Harbor, Keweenaw County

      Hydraena Kugelann, 1794*: 579
        Hydraena angulicollis Notman, 1921*: 146 - Perkins 1980: 81-83, 443
        Hydraena pennsylvanica Kiessenwetter, 1849*: 166 - Perkins 1980: 81, 96-98, 443; neootype from Bryant Road, Emmet County; record from Grand Ledge, Eaton Co. as H. atlantica (Perkins 1980)
          (syn.) Hydraena atlantica Perkins, 1980*: 488

      Ochthebius Leach, 1815*: 95
        Ochthebius (Asiobates) cribricollis LeConte, 1850*: 217 - Perkins 1980: 386, 388-389, lectotype from Eagle Harbor, Keweenaw County
        Ochthebius (Ochtebius) kaszabi Janssens, 1967*: 56 - MN, ON (may be found in Isle Royale)(Perkins 1980: 282, 325-328)
        Ochthebius (Ochtebius) lineatus LeConte, 1852*: 211 - WI, western, but one record from northern WI, expect in w UP (Perkins 1980:314-318, 323)
          (syn.) Ochthebius milleri Hatch, 1965: 18

      References

        Hatch MH. 1965. The beetles of the Pacific Northwest. Part IV: Macrodactyles, Palpicornes, and Heteromera. University of Washington Press, Seattle, 268 pages.
        Janssens E. 1967. 91. Hydraenidae. Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei (Coleoptera). Reichenbachia 9(5): 53-58.
        Kugelann JG .1794. Verzeichniß der in einigen Gegenden Preußens bis jetzt entdeckten Käfer-Arten, nebst kurzen Nachrichten von denselben, pp. 513-582 in Schneider DH (ed.), Neuestes Magazin für die Liebhaber der Entomologie. Vol. I, part 5. D. H. Schneider, Stralsund.
        Kiesenwetter EAH, von. 1849. Monographische revision der Gattung Hydraena. Linnaea Entomologica 4: 156-190, 425-427.
        Leach WE. 1815. Entomology, pp. 57-172 in Edinburgh Encyclopedia, Volume 9(1)(Brewster, ed.). Edinburgh. (VIEW)
        LeConte JL. 1852. Descriptions of new Coleoptera from California. Annals of the Lyceum of Natural History of New York 5: 185-216.
        LeConte JL. 1850. General remarks upon the Coleoptera of Lake Superior, pp 201-242 in Agassiz L, LeConte JL, Lake Superior: Its physical character, vegetation, and animals, compared with those of other and similar regions. Gould, Kendall and Lincoln, Boston, 428 pp., illus.
        Mulsant ME. 1844. Histoire naturelle des coléoptères de France. Vol. 3. Palpicornes, Lyon. (PDF)
        Notman H. 1921. Some new genera and species of Coleoptera collected at Westfield, Chautauqua Co., N.Y. Journal of the New York Entomological Society 24: 146-147.
        Orchymont A, d'. 1943b. Faune du nord-est bresilien (recoltes du Dr. O. Schubart). Palpicornia. Mémoires du Musée royal d'histoire naturelle de Belgique, deuxieme serie, 28: 1-85.
        Perkins PD. 1980. Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae 16: 3-554.

    HYDROPHILIDAE Latreille, 1802 (Water Scavenger Beetles)
    The information contained on this list is tentative, based on few observations of museum specimens (currently being assessed) and a limited literature search (Downie and Arnett 1996, Willson 1970). Terrestrial species of Sphaeridiinae, including those in the genera Cercyon, Cryptopleurum, Paraceryon, Pemelus, Phaenonotum, Sphaeridium, and Tectosternum are not included, as they are associated with decaying plant or fungal matter, and animal dung, and are only rarely (or incidentally) collected in aquatic systems. A number of species of Cercyon and Phaenonotum are eith semi- or peri-aquatic species, and associated with aquatic habitats, and these are included. Consult Smetana (1988) or Downie and Arnett (1996) for more information.

      Anacaena Thomson, 1859 (Subfamily Hydrobiinae, Tribe Hydrobiini)
        Anacaena lutescens Stephens, 1829 - (as A. limbata (Fabricius, 1792), see Hilsenhoff 1995c), UMMZ; see also Willson 1970:80-81; Smetana 1988:87-88

      Berosus Leach, 1817 (Subfamily Berosinae)
        Berosus aculeatus LeConte, 1855 - Smetana 1988:51; Downie and Arnett 1996
        Berosus fraternus LeConte, 1855 - Downie and Arnett 1996, NMNH (web page database); see also Smetana 1988:54-55
          (syn.) Berosus californicus Motschulsky, 1859
        Berosus hatchi Miller, 1965 - ON, WI (Smetana 1988:53-54)
        Berosus infuscatus LeConte, 1855 - Charity Island (44.030°N, 83.437°W), specimens in UMMZ, NMNH (web page database)
        Berosus ordinatus LeConte - IN, MN (and eastward)
        Berosus pantherinus LeConte, 1855 - IL, IN, OH, ON, WI
        Berosus peregrinus (Herbst, 1797) Hydrophilus (orig.) - UMMZ; see also Smetana 1988:50
          (syn.) Berosus auritus Melsheimer, 1846
          (syn.) Berosus peregrinus, LeConte 1855, Matta, 1974
          (syn.) Berosus subsignatus LeConte, 1855
          (syn.) Berosus immaculatus Zimmermann, 1869
        Berosus striatus (Say, 1825) Hydrophilus (orig.) - UMMZ, NMNH (web page database); see also Smetana 1988:51-52
        Berosus stylifer Horn, 1873 - Smetana 1988:55-56
          (syn.) Berosus falcatus Sharp, 1882

      Cercyon Leach, 1817 (Subfamily Spaheridiinae, Tribe Cercyonini)
        Cercyon (Prostercyon) roseni Knisch, 1922 - Smetana 1988:136-137 (peri- to semi-aquatic, along wetlands, ponds, bogs and lakes)
        Cercyon (Cercyon) connivens Fall, 1924 - Smetana 1988: 141 (peri-aquatic)
        Cercyon (Cercyon) herceus Smetana, 1978 - Smetana 1988: 142-143 (peri-aquatic)
          (syn.) Cercyon tristis, Horn 1890 (nec Illiger, 1801)
          (syn.) Cercyon herceus Smetana, 1978; plus spp. herceus and frigidus
        Cercyon (Cercyon) erraticus Smetana, 1978 - Smetana 1988: 144 (peri-aquatic)
        Cercyon (Cercyon) mendax Smetana, 1978 - Smetana 1988: 145 (peri-aquatic)
        Cercyon (Cercyon) praetextatus (Say, 1825) Sphaeridium (orig.) - Smetana 1988: 145-146
          (syn.) Cercyon testaceus Blatchley, 1923
        Cercyon (Cercyon) marinus Thomson, 1853 - Smetana 1988: 147-148 (semi-aquatic)
        Cercyon (Cercyon) cinctus Smetana, 1978 - Smetana 1988: 148 (peri-aquatic, similar habitat preferences with marinus)

      Chaetarthria Stephens, 1833 (Subfamily Chaetarthriinae)
        Chaetarthria atra (LeConte, 1863) Cyllidium (orig.) - IL, IN (Smetana 1988: 57-58)
          (syn.) Chaetarthria nigrella, Wickham 1895 (nec LeConte, 1861)
        Chaetarthria pallida (LeConte, 1861) Cyllidium (orig.) - Cheboygan Co., UMMZ, NMNH (web page database); see also Smetana 1988: 58
          (syn.) Cyllidum nigriceps LeConte, 1861

      Crenitis Bedel, 1881 (Subfamily Hydrobiinae)
        Crenitis digesta (LeConte, 1855) Hydrobius (orig.) - Willson 1970: 78-79; see also Smetana 1988: 81-82)
          (syn.) Paracymus digestus Winters, 1926
          (syn.) Crenophilus digestus d'Orchymont, 1942

      Cymbiodyta Bedel, 1881 (Subfamily Hydrobiinae, Tribe Helocharini)
        Cymbiodyta acuminata Fall, 1924 - UMMZ; Willson 1970: 52-53; Smetana 1974: 68-70; see also Smetana 1988: 110-111
        Cymbiodyta blanchardi Horn, 1890 - UMMZ; Willson 1970: 53; see also Smetana 1974: 61-64; Smetana 1988: 107-108
        Cymbiodyta chamberlaini Smetana, 1974 - UMMZ; Smetana 1974: 57-61; Smetana 1988: 106-107
        Cymbiodyta minima Notman, 1919 - UMMZ, NMNH (web page database); Willson 1970: 54; Smetana 1974: 73-76; see also Smetana 1988: 111-112
          (syn.) ?Hydrobius marginellus Kirby, 1837, in Richardson
          (syn.) Philidrus lacustris Horn, 1873
          (syn.) Cymbiodyta lacustris Wickham, 1895
          (syn.) Cymbiodyta acuminata Hatch, 1933 (misid).
        Cymbiodyta semistriata (Zimmermann, 1869) Philhydrus (orig.) - UMMZ, NMNH (web page database); Smetana 1974: 48-51; see also Smetana 1988: 103, 105
          (syn.) Philhydrus fimbriatus LeConte, 1855
          (syn.) Philidrus fimbriatus Provancher, 1877
          (syn.) Cymbiodyta fimbriata, Horn 1890 and authors Cymbiodyta toddi Spangler, 1966 - Willson 1970: 56; Smetana 1974: 51-54; see also Smetana 1988: 104, 106
          Cymbiodyta vindicata Fall, 1924 - UMMZ, NMNH (web page database); Willson 1970: 55-56; Smetana 1974: 64-67; see also Smetana 1988: 108-109

        Dibolocelus Bedel, 1892 (Subfamily Hydrophilinae)
          Dibolocelus ovatus (Gemminger & Harold, 1868) Hydrophilus (orig.) - Willson 1970; see also Smetana 1988: 124
            (syn.) Hydrophilus ovilis Ziegler, 1844 (nec Laport, 1840)

        Enochrus Thomson, 1859 (Subfamily Hydrobiinae, Tribe Helocharini)
          Enochrus (Methydrus) blatchleyi (Fall, 1924) Philhydrus (orig.) - Willson 1970: 62; see also Smetana 1988: 92
          Enochrus (Methydrus) cinctus (Say, 1824) Hydrophilus - Willson 1970: 63-64; record at Albertson College of Idaho Collection; see also Smetana 1988: 92-93
            (syn.) Philhydrus limbalis Melsheimer, 1846
          Enochrus collinus (Fall, 1924) Philhydrus (orig.) - WI
          Enochrus (Methydrus) consors LeConte, 1863 Philhydrus (orig.) - Willson 1970: 65; see also Smetana 1988: 93)
          Enochrus (Methydrus) consortus Green, 1946 - Willson 1970: 64-65; see also Smetana 1988: 93-94
          Enochrus (Lumetus) diffusus (LeConte, 1855) Philhydrus (orig.) - record at Albertson College of Idaho Collection; western species with isolated eastern populations (Smetana 1988: 99); see also Willson 1970: 66-67
          Enochrus (Lumetus) hamiltoni (Horn, 1890) Philhydrus (orig.) - Downie and Arnett 1996; see also Smetana 1988: 99-100
            (syn.) Philhydrus lividus Walker, 1866
            (syn.) Philhydrus conjunctus Fall, 1901
            (syn.) Enochrus collinus Brown, 1931
            (syn.) Enochrus horni Leech, 1950
            (syn.) Enochrus pacificus Leech, 1950
            (syn.) Enochrus pyretus Leech, 1950
          Enochrus (Methydrus) ochraceus (Melsheimer, 1846) Philhydrus (orig.) - Willson 1970: 62-63; record at Albertson College of Idaho Collection; see also Smetana 1988: 95
            (syn.) Philhydrus lacustris LeConte, 1855
            (syn.) Philhydrus simplex LeConte, 1863
          Enochrus (Methydrus) perplexus (LeConte, 1855) Philhydrus (orig.) - Willson 1970: 54-55 (as C. fimbriata), 65-66; see also Smetana 1988: 94
            (syn.) Cymbiodyta fimbriatua (Melsheimer, 1846)
          Enochrus (Methydrus) pygmaeus nebulosus (Say, 1824) Hydrophilus - Willson 1970: 60-61; see also Smetana 1988: 96
            (syn.) Hydrophilus pygmaeus Fabricius, 1792
          Enochrus (Methydrus) sayi Gunderson, 1977 - Downie and Arnett 1996; Smetana 1988: 97

        Helocombus Horn, 1890 (Subfamily Hydrobiinae, Tribe Helocharini)
          Helocombus bifidus (LeConte, 1855) Philhydrus (orig.) - Willson 1970: 50-51; see also Smetana 1988: 113

        Helophorus Fabricius, 1775 (Subfamily Helophorinae)
          Helophorus (Rhopalelophorus) angusticollis d'Orchymont - Downie and Arnett 1996; see also Smetana 1988: 31, 43-44
          Helophorus (Rhopalelophorus) artus Smetana - IL
          Helophorus (Rhopalelophorus) furius Smetana, 1987 - ON, WI (northern: Smetana 1988: 27-28)
            (syn.) Helophorus frater Smetana, 1985 (nec d'Orchymont, 1926)
          Helophorus (Helophorus) grandis Illiger, 1798 Elophorus (orig.) - record from Huron Co.; see Smetana 1988: 20-21 (introduced species from Europe)
            (syn.) Helophorus aquaticus, Darlington 1927, Brown 1940 (nec Linné 1758)
          Helophorus (Rhopalelophorus) inflectus McCorkle, 1965 Elophorus (orig.) - Smetana 1988: 25-26; Downie and Arnett 1996
          Helophorus (Rhopalelophorus) lacustris LeConte, 1850 - Downie and Arnett 1996; see also Smetana 1988: 39-40
          Helophorus (Rhopalelophorus) latipenis - WI (syn. or ssp.?)
          Helophorus (Rhopalelophorus) linearis LeConte, 1850 - IL, IN (Waltz 2002: 178), OH, ON, WI (Smetana 1988: 36-37)
            (syn.) Helophorus pallens Casey, 1884
            (syn.) Helophorus oblitus Sharp, 1915
          Helophorus (Rhopalelophorus) lineatus Say, 1823 - Downie and Arnett 1996; see also Smetana 1988: 29-30
            (syn.) Helophorus ventralis Motschulsky, 1860
            (syn.) Helophorus obsoletesulcatus Motschulsky, 1860
          Helophorus (Rhopalelophorus) marginicollis Smetana, 1985 - Smetana 1988: 44; Downie and Arnett 1996
          Helophorus (Rhopalelophorus) nitiduloides d'Orchymont, 1945 - Downie and Arnett 1996; see also Smetana 1988: 37-38
          Helophorus (Rhopalelophorus) nitidulus LeConte, 1855 - Smetana 1988: 28-29; Downie and Arnett 1996
            (syn.) Helophorus gregalis d'Orchymont, 1945
          Helophorus (Rhopalelophorus) oblongus LeConte, 1850 - Downie and Arnett 1996; see also Smetana 1988: 24-25
          Helophorus (Rhopalelophorus) orchymonti Smetana, 1985 - ON, WI (Smetana 1988: 43)
          Helophorus (Rhopalelophorus) orientalis Motschulsky, 1860 - Downie and Arnett 1996; see also Smetana 1988: 34-35
            (syn.) Helophorus brevipalpis, McCorkle, 1965 (nec Bedel, 1881)
          Helophorus (Rhopalelophorus) sempervarians Angus, 1970 - Downie and Arnett 1996; see also Smetana 1988: 32-33
            (syn.) Elophorus jacutus, McCorkle, 1965 (nec Poppius, 1907)
            (syn.) Helophorus jacutus, d'Orchymont, 1945 (nec Poppius, 1907)
          Helophorus (Cyphelophorus) tuberculatus Gyllenhal - Downie and Arnett 1996; see also Smetana 1988: 45-46

        Hydrobius Leach, 1815 (Subfamily Hydrobiinae, Tribe Hydrobiini)
          Hydrobius fuscipes (Linnaeus, 1758) Dytiscus (orig.) - Willson 1970: 69, NMNH (web page database), record at Albertson College of Idaho Collection; see also Smetana 1988: 69-70
          Hydrobius melaenus (Germar, 1824*) Sphaeridium (orig.) - Willson 1970: 69-71 (as H. globosus); see also Smetana 1988: 70-71
            (syn.) Hydrophilus globosus Say, 1824
            (syn.) Hydrobius globosus LeConte, 1855
            (syn.) Hydrobius globulosus Provancher, 1877
            (syn.) Hydrobius melaenus Winters, 1926
          Hydrobius tumidus LeConte, 1855 - IN

        Hydrochara Berthold, 1827 (Subfamily Hydrophilinae)
          Hydrochara leechi Smetana, 1980 - IL, OH, WI, may occur in s LP (Smetana 1980: 38-41)
          Hydrochara obtusata (Say, 1823) Hydrophilus - Willson 1970: 22-23; NMNH (web page database); Smetana 1980: 24-30; see also Smetana 1988: 127
          Hydrochara simula Hilsenhoff & Tracy - Hilsenhoff and Tracy 1982: 101; see also Smetana 1988: 128
          Hydrochara soror Smetana, 1980 - Smetana 1980: 30-34; see also Smetana 1988: 128
          Hydrochara spangleri Smetana, 1980 - IL, IN, OH, WI, may occur in sw LP (Smetana 1980: 34-36)

        Hydrochus Leach, 1817 (Subfamily Hydrochinae)
          Hydrochus brevitarsis Knisch, 1922 - Smetana 1988: 12; Hilsenhoff 1995: 42
          Hydrochus currani Brown, 1929 - ON, WI (Smetana 1988: 11-12)
          Hydrochus granulatus Blatchley - IN, ON, WI
          Hydrochus excavatum LeConte - IN
          Hydrochus granulatus Blatchly, 1910 - QB-BC, NY-MN (Smetana 1988: 17-18)
            (syn.) Hydrochus ouelleti Leech, 1848
          Hydrochus inaequalis LeConte - IN
          Hydrochus neosquamifer Smetana, 1988 - ON, WI (Smetana 1988: 14-15)
          Hydrochus pseudosquamifer Miller, 1965 - IN, ON, WI (Smetana 1988: 15)
          Hydrochus rufipes Melsheimer - IL, IN, OH, WI (Smetana 1988: 18)
            (syn.) Hydrochus impressus Zimmermann, 1868
          Hydrochus rugosus Mulsant - IN
          Hydrochus scabratus Mulsant, 1844 - Downie and Arnett 1996; see also Smetana 1988: 11
            (syn.) Hydrochus gibbosus Melsheimer, 1846
          Hydrochus squamifer LeConte, 1855 - IN, ON, WI (Smetana 1988: 13-14)
            (syn.) Hydrochus laticollis Notman, 1919
          Hydrochus subcupreous Randall, 1838 - IN, WI (Smetana 1988: 15-17)

        Hydrophilus Geoffroy, 1762 (Subfamily Hydrophilinae)
          Hydrophilus triangularis Say, 1823 - Willson 1970: 24-25; see also Smetana 1988: 123-124

        Laccobius Erichson, 1837 (Subfamily Hydrobiinae, Tribe Hydrobiini)
          Laccobius agilis Randall, 1838 Hydrophilus (orig.) - Willson 1970: 38-39; see also Smetana 1988: 61-62
            (syn.) Laccobius ellipticus, Stace Smith 1930 (nec LeConte, 1855)
            (syn.) Laccobius agilis Gentili, 1985
          Laccobius carri d'Orchymont, 1942 - ON, may occur in UP (Smetana 1988: 67-68)
          Laccobius cinereus columbianus Miller, 1965 - Downie and Arnett 1996; see also Smetana 1988: 64-65
          Laccobius fuscipunctatus Hilsenhoff, 1995 - WI (Hilsenhoff 1995c: 112-114)
          Laccobius minutoides d'Orchymont, 1942 - Willson 1970: 39-40; see also Smetana 1988: 62-63
          Laccobius reflexipennis Cheary, 1971 - ON, WI (Smetana 1988: 66-67)
            (syn.) Laccobius reflexipennis Malcolm, 1979
          Laccobius spangleri Cheary, 1971 - Willson 1970: 40-44; Cheary 1971; NMNH (web page database); see also Smetana 1988: 67
            (inv.) Laccobius spangleri Willson, 1970, manuscript, never published
            (syn.) Laccobius spangleri Malcolm, 1979
          Laccobius teneralis Cheary, 1971 - ON, may occur in s. LP (Smetana 1988: 65-66)
          Laccobius truncatipenis Miller, 1965 - Willson 1970: 44-48 (as L. arenarius); Cheary 1971 (as L. arenarius); NMNH (web page database); see also Smetana 1988: 68-69
            (syn.) Laccobius arenarius Willson 1970, unpublished MS thesis
            (syn.) Laccobius arenarius Cheary, 1971

        Paracymus Thomson, 1867 (Subfamily Hydrobiinae, Tribe Hydrobiini)
          Paracymus confluens Wooldridge, 1966 - Wooldridge 1966; Willson 1970: 74-75; Smetana 1988: 75-76
          Paracymus confusus Wooldridge, 1966 - Downie and Arnett 1996; see also Willson 1970: 75; Smetana 1988: 77
          Paracymus despectus (LeConte, 1863) Hydrobius - Willson 1970: 73-74; NMNH (web page database); see also Smetana 1988: 78
          Paracymus subcupreus (Say, 1825) Hydrobius - Willson 1970: 75-76, NMNH (web page database); see also Smetana 1988: 76-77

        Phaenonotum Sharp, 1882 (Subfamily Spaheridiinae, Tribe Sphaeridiini)
          Phaenonotum exstriatum (Say, 1835) Hydrophilus (orig.) - Willson 1970: 13-14; Smetana 1988: 130 (often semi-aquatic, at edges of wet habitats and water, wet moss and Sphagnum)
            (comb.)Cyclonotum estriatum LeConte, 1855 (unjustified emendation)
            (syn.) Cyclonotum semiglobosum Zimmermann, 1869
            (syn.) Phaenonotum dubium Sharp, 1882
            (spell.) Phaenonotum estriatum (Say, 1835), Willson 1970: 13

        Sperchopsis LeConte, 1862 (Subfamily Hydrobiinae, Tribe Hydrobiini)
          Sperchopsis tessellatus (Ziegler, 1844) Spercheus (orig.) - Willson 1970: 71-72; record at Albertson College of Idaho Collection; Bright coll.; see also Smetana 1988: 72
            (syn.) Sperchopsis tessellatus LeConte, 1861
            (syn.) Hydrobius tessellatus Horn, 1873
            (syn.) Hydrocyclus tessellatus Winters, 1926
            (syn.) Sperchopsis tessellatus Spangler, 1961

        Tropisternus Solier, 1834 (Subfamily Hydrophilinae)
          Tropisternus blatchleyi modestus d'Orchymont, 1938 - Willson 1970: 32 (as T. modestus); see also Smetana 1988: 119-120
            (syn.) Tropisternus modestus d'Orchymont, 1938
          Tropisternus columbianus Brown, 1931 - Willson 1970: 29-30; see also Smetana 1988: 122
          Tropisternus ellipticus LeConte, 1855 Hydrophilus - IL, WI (Smetana 1988: 118-119; see also Willson 1970: 29)
            (syn.) Tropisternus affinis Motschulsky, 1859
            (syn.) Tropisternus concolor Sharp, 1882
          Tropisternus glaber (Herbst, 1797) Hydrophilus (orig.) - Willson 1970: 33-34; Ryker 1972; see also Smetana 1988: 121
          Tropisternus lateralis nimbatus (Say, 1823) Hydrophilus (orig.) - Willson 1970: 28; Ryker 1972 (as T. nimbatus); see also Smetana 1988: 118
            (syn.) Tropisternus nimbatus Summers, 1874
            (syn.) Tropisternus lateralis Horn, 1876; LaRivers 1949
            (syn.) Tropisternus lateralis nimbatus Young, 1954
          Tropisternus mixtus (LeConte, 1855) Hydrophilus (orig.) - Willson 1970: 30-31; Ryker 1972; see also Smetana 1988: 119
          Tropisternus natator d'Orchymont, 1938 - Willson 1970: 32-33; Ryker 1972; see also Smetana 1988: 120-121
            (syn.) Tropisternus glaber, LeConte, 1859; Wooldridge, 1967
            (syn.) Hydrophilus glaber, LeConte, 1855 (nec Herbst, 1797)

    LAMPYRIDAE Latreille, 1817 (Firefly Beetles)

    Several taxa of these fascinating beetles have larvae known to live in the air spaces of aquatic plants, or are associated with aquatic or semiaquatic plants in aquatic habitats. A number of species have larvae that are adapted to feed on aquatic snails, and perhaps annelids; some larvae of Asian species have tracheal gills that permit a truly aquatic existence. There are 6 species in 2 genera that have been recorded in Michigan, with another 3 species in 2 genera likely to be found based on their regional distribution. This list is based on information kindly provided by Dr. Jim Lloyd (pers.comm. 2003), and records listed in Downie and Arnett (1996).

      Photinus LaPorte, 1833

        Photinus ardens LeConte, 1851 - Downie and Arnett 1996; Lloyd pers.comm. (Marquette, Delta, Chippewa, Charlevoix, Cheboygan, Roscommon, Huron, St. Clair, Washtenaw)
        Photinus consimilis Green (complex), 1956 - Downie and Arnett 1996; Lloyd pers.comm., as consimilis-complex (Schoolcraft, Chippewa, Oceana, Livingston, Oakland, St. Clair)
        Photinus obscurellus LeConte, 1851 - Downie and Arnett 1996; Lloyd pers.comm (Houghton, Iron, Dickinson, Marquette, Mackinaw, Chippewa, Berrien, St. Joseph, Washtenaw, Wayne, St. Clair)

      Pyractomena LeConte, 1850

        Pyractomena angulata (Say, 1825) Lampyris (orig.) - e. USA, Canada (Lloyd, pers.comm.; Luk et al. 2011: 68, 72)
        Pyractomena borealis (Randall, 18258 Lampyris (orig.) - OH, e. USA, Canada (Archangelsky and Branham 1998; Lloyd, pers.comm.; Luk et al. 2011: 69, 73)
        Pyractomena dispersa Green, 1958 - Downie and Arnett 1996; Lloyd pers. comm. (Kent, Washtenaw, Wayne)
        Pyractomena linearis LeConte, 1851 - Downie and Arnett 1996; Lloyd pers.comm. (Marquette, Dickinson, Mackinac, Livingston, Washtenaw)
        Pyractomena lucifera Melsheimer, 1846 - Downie and Arnett 1996; Lloyd pers.comm. (Midland, Clinton, Livingston, Washtenaw, Oakland)
        Pyractomena sinuata Green, 1958 - IL, MN, WI (Lloyd pers.comm.)

      Pyropyga Motschulsky, 1852

        Pyropyga decipiens (Harris, 1836) Lampyris (orig.) - Lloyd pers.comm. (Keweenaw, widespread in LP)
        Pyropyga nigricans (Say, 1823) Lampyris (orig.) - Lloyd 1999; Lloyd pers.comm. (widespread in state)

    LUTROCHIDAE Kasap & Crowson, 1975 (Long-toed or Travertine Beetles) (Polyphaga)
    Lutrochus luteus adult - from Brown 1972

    These beetles share the same habitats as that of dryopid and elmid riffle beetles (Dryopidae, Elmidae) and water-pennies (Psephenidae), although there is a preference for calcarious deposits from mineral springs (Brown 1972) (or possibly areas with high rates of instream photosynthetic activity, when calcium carbonate precipitates). There is one species that reaches our area. In Wisconsin, larvae were found in rocks, and adults were found in various lotic habitats of mixed substrate, with ovipositing adults occuring on wood and the travertine of rocks in shallow riffles (Hilsenhoff and Schmude 1992).

      Lutrochus Erichson, 1847
        Lutrochus laticeps Casey, 1893 - Brown 1972

    PSEPHENIDAE Lacordaire, 1854 (Water Pennies)

    Larva of Psephenus herricki (Copyright © 2006 Ethan Bright)
    Psephenus herricki - larva - copyright 2006 Ethan Bright

      Ectopria LeConte, 1853 (Subfamily Eubriinae)
        Ectopria leechi Brigham, 1981
        Ectopria nervosa (Melsheimer, 1845) Eubria (orig.) - Yankoviak and McCafferty 1996: 205
          (syn.) Eubria thoracica Ziegler, 1845
          (syn.) Ectopria tarsalis LeConte, 1853
          (syn.) Ectopria tibialis LeConte, 1853

      Psephenus Haldeman 1853 (Subfamily Psepheninae)
        Psephenus herricki (De Kay, 1844) Fluvicola (orig., described as an isopod) - Brown and Murvosh 1974: 315-322; widespread in state
          (nomen nudum) Eurypalpus lecontei De Jean, 1837
          (syn.) Psephenus lecontei Haldeman, 1853

    PTILODACTYLIDAE Laporte, 1836 (Toad-Winged Beetles)

    Larvae are known to live in semiaquatic or aquatic habitats, such as decaying damp logs and organic muck. Recently the larva of Paralichas (=Odontyonyx) trivattis has been described, reared from specimens sampled from spring seeps in eastern Pennsylvania (Funk and Fenstermacher 2002). Downie and Arnett (1996) list a questionable record from Indiana, and since the biology and distribution of this species are not well known, I have included this taxa as it conceivably may occur in our area, especially in morainal/outwash areas of southern Michigan where groundwater seeps occur.

      Odontonyx Guérin-Méneville, 1843

        Odontonyx trivittis (Germar, 1824*) Dasytes - IN?, PA

    SCIRTIDAE Fleming, 1821 (=Helodidae) (Marsh Beetles)

      Cyphon Paykull, 1799
        Cyphon collaris (Guérin-Meneville, 1843) Elodes (orig.) - IN, ON
        Cyphon confusus Brown, 1930 - Young and Stribling 1990: 201-202
          (syn.) Cyphon horioni Klausnitzer, 1976
        Cyphon drymophilous Young & Stribling, 1990 - OH, WI (Young and Stribling 1990: 202-203)
        Cyphon obscurus (Guérin-Meneville, 1843) Elodes (orig.) - IN, OH, ON
        Cyphon padi (Linnaeus, 1758) Chrysomela (orig.) - IN, ON
        Cyphon ruficollis (Say, 1825) Lampyris (orig.) - IN (widespread east and central)
        Cyphon variabilis (Thunberg, 1785) Cantharis - Downie and Arnett 1996

      Elodes Latreille, 1796

        Elodes fuscipennis Guérin-Meneville 1843 - IN
        Elodes maculicollis (Horn, 1880) Helodes (orig.) - ON
        Elodes pulchella Guérin-Meneville, 1843 - IN
        Elodes thoracica Guérin-Meneville, 1843 - IN, ON

      Microcara Thomson 1859

        Microcara explanata LeConte, 1866 Helodes (orig.) - Downie and Arnett 1996, shores of Lake Michigan

      Prionocyphon Redtenbacher, 1858

        Prionocyphon discoideus (Say, 1825) Cyphon (orig.) - Downie and Arnett 1996

      Scirtes Illiger, 1807

        Scirtes orbiculatus (Fabricius, 1801) Cyphon (orig.) - IN, OH
        Scirtes tibialis (Guérin-Meneville, 1843) Scirtes (orig.) - IN, ON, WI

    STAPHYLINIDAE Latreille, 1802 (Rove Beetles)

    This large family of slender beetles are characteristic in having short, truncated elytra that leaves several abdominal segments dorsally visible. Most species are terrestrial, but a few are associated with aquatic environments, principally with shore-line environments and the surface of littoral habitats. Psephidonus is common under stones at or near the edge of running water. Carpelimus and Thinobius is generally found along littoral areas of lakes and rivers among leaf litter. Not all species of Bledius and Stenus are semiaquatic, and provisionally terrestrial taxa are included in the list based on citations in Downie and Arnett (1996); museum and other collection specimens have have only begun to be examined.

      Bledius Leach, 1819 (Subfamily Osytelinae)

        Bledius analis LeConte, 1863 - Downie and Arnett 1996
        Bledius annularis LeConte, 1863 - IN (widespread)
        Bledius armatus Say - UMMZ Insect Collection (syn. of what name?)
        Bledius assimilis Casey, 1889 - Downie and Arnett 1996
        Bledius confusus LeConte, 1877 - Downie and Arnett 1996
        Bledius emarginatus (Say, 1834) Oxytelus - Downie and Arnett 1996
        Bledius fumatus LeConte, 1863 - Downie and Arnett 1996
        Bledius gravidus Casey, 1889 - Downie and Arnett 1996
        Bledius ineptus Casey, 1889 - Downie and Arnett 1996
        Bledius nitidicollis LeConte, 1863 - IL, OH (and northwest)
        Bledius omega Herman, 1983 - Downie and Arnett 1996
        Bledius pallipennis (Say, 1823) Oxytelus - IL, OH
        Bledius rotundicollis LeConte, 1877 - IL, MN, WI
        Bledius ruficornis LeConte, 1863 - Downie and Arnett 1996
        Bledius semiferrugineus LeConte, 1863 - UMMZ Insect Collection
        Bledius sinuatus LeConte, 1877 - IL, ON
        Bledius suturalis LeConte, 1863 - ON (widespread)
        Bledius tarandus Herman, 1970 - ON, WI
          (syn.) Bledius borealis Blatchley
        Bledius tau LeConte, 1877 - Downie and Arnett 1996
        Bledius trugidus Casey, 1889 - Downie and Arnett 1996
        Bledius viriosus Herman, 1983 - ON, WI

      Carpelimus Leach, 1819 (Subfamily Osytelinae)

        Carpelimus bilineatus (Stephen), 1832 Carpalimnus [sic] - IN
        Carpelimus delicatus (Casey, 1889) Trogophloeus - Downie and Arnett 1996
        Carpelimus fallax (Casey, 1889) Trogophloeus - Downie and Arnett 1996
        Carpelimus morio (Erichson, 1840) Trogophloeus - Downie and Arnett 1996
        Carpelimus quadripunctatus (Say, 1834) Stenus - IN, OH

      Psephidonus Gistel, 1856 (=Geodromicus Redtenbacher) (Subfamily Omaliinae)

        Psephidonus brunneous (Say, 1823) Anthophagus - UMMZ Insect Division
        Psephidonus strictus (Fauvel), 1889 - Downie and Arnett 1996
        Psephidonus ovipennis (LeConte, 1878) Geodromicus - widespread, disjunct (New York, Utah)

      Stenus Latreille, 1796 (Subfamily Steninae)

        Stenus advenus (Casey, 1884) Areus - Downie and Arnett 1996
        Stenus alacer Casey, 1884 - IL, IN
        Stenus argus Gravenhorst, 1806 - IN
        Stenus angustus Casey, 1884 - Downie and Arnett 1996
          (syn.) Stenus indigens Casey, 1884
          (syn.) Stenus pettiti Casey, 1884
          (syn.) Stenus placidus Casey, 1884
          (syn.) Stenus mendax Casey, 1884
        Stenus annularis Erichson, 1840 - Downie and Arnett 1996
        Stenus brivioi Puthz, 1972 - Downie and Arnett 1996
        Stenus callosus Erichson, 1840 - ? (no locality in Downie and Arnett 1996)
        Stenus canaliculatus Gyllenhal, 1827 - Downie and Arnett 1996
        Stenus capucinus Boheman, 1858 - Downie and Arnett 1996
          (syn.) Stenus varipes Casey, 1884
        Stenus carinicollis Casey, 1884 - Downie and Arnett 1996
        Stenus caseyi Puthz, 1971 - Downie and Arnett 1996
        Stenus colonus Erichson, 1840 - Downie and Arnett 1996
          (syn.) Stenus militaris Casey, 1884
          (syn.) Stenus suspectus Blatchley
          (syn.) Stenus obstrusus Casey, 1884
          (syn.) Stenus integer Casey, 1884
        Stenus comma LeConte, 1863 - Downie and Arnett 1996
        Stenus colon Say, 1834 - Downie and Arnett 1996
        Stenus croceatus (Casey, 1884) - Downie and Arnett 1996
        Stenus curtus Casey, 1884 - Downie and Arnett 1996
        Stenus dissentiens (Casey, 1884) - Downie and Arnett 1996
          (syn.) Stenus hubbardi (Casey, 1884)
        Stenus egenulus Puthz - Downie and Arnett 1996
        Stenus egenus Erichson, 1840 - IN (widespread)
        Stenus erythropus Melsheimer, 1844 - Downie and Arnett 1996
        Stenus femoratus Say, 1834 - Downie and Arnett 1996
        Stenus flavicornis Erichson, 1840 - Downie and Arnett 1996
        Stenus fulvoguttatus Notman, 1920 - Downie and Arnett 1996
        Stenus gratiosus Casey, 1884 - Downie and Arnett 1996
          (syn.) Stenus hirsutus Casey, 1884
        Stenus immarginatus Maklin, 1853 - widespread east
          (syn.) Stenus monticola Casey, 1884
        Stenus insperatus Puthz, 1972 - Downie and Arnett 1996
        Stenus intrusus Casey, 1884 - Downie and Arnett 1996
        Stenus juno Fabricius, 1801 - Downie and Arnett 1996
        Stenus laccophilus Casey, 1884 - UMMZ insect collection
          (syn.) Stenus edax Notman
          (syn.) Stenus rugifer Casey, 1884
        Stenus mammops Casey, 1884 - Downie and Arnett 1996
        Stenus morio Gravenhorst, 1806 - Downie and Arnett 1996
          (syn.) Stenus morio Gravenhorst, 1806
        Stenus nanus Stephen, 1832 - Downie and Arnett 1996
        Stenus neglectus Casey, 1884 - Downie and Arnett 1996
        Stenus parallelopipedus Casey, 1884 - Downie and Arnett 1996
          (syn.) Stenus animatus Casey, 1884
        Stenus pauperculus Casey, 1884 - Downie and Arnett 1996
        Stenus pubescens Stephens, 1832 - Downie and Arnett 1996 (native of Palearctic?)
        Stenus pudicus Casey, 1884 - Downie and Arnett 1996
          (syn.) Stenus juvencus Casey, 1884
        Stenus pumilio Erichson, 1839 - Downie and Arnett 1996
        Stenus punctatus Erichson, 1840 - Downie and Arnett 1996
        Stenus pusillimus Puthz, 1971 - Downie and Arnett 1996
        Stenus reconditus (Casey, 1884) - Downie and Arnett 1996
          (syn.) Stenus canadensis (Casey, 1884)
        Stenus rossi Sanderson, 1957 - Downie and Arnett 1996
        Stenus scrupens Casey, 1884 - Downie and Arnett 1996
        Stenus semicolon LeConte, 1863 - Downie and Arnett 1996
        Stenus strangulatus Casey, 1884 - Downie and Arnett 1996
        Stenus scabiosus Casey, 1884 - Downie and Arnett 1996
        Stenus sibiricus Sahlberg, 1880 - Downie and Arnett 1996
          (syn.) Stenus deductor Casey, 1884
          (syn.) Stenus montanus Casey, 1884
        Stenus stygicus Say, 1834 - Downie and Arnett 1996
        Stenus tenuis Casey, 1884 - Downie and Arnett 1996
          (syn.) Stenus pauper Casey, 1884
          (syn.) Stenus subtilis Casey, 1884
          (syn.) Stenus ingratus Casey, 1884
        Stenus varipes Casey, 1884 - Downie and Arnett 1996
        Stenus vicinus Casey, 1884 - Downie and Arnett 1996
        Stenus vinnulus Casey, 1884 - Downie and Arnett 1996

      Thinobius Kiesenwetter, 1844 (Subfamily Osytelinae)

        Thinobius brachypterusLeConte, 1877 - Downie and Arnett 1996
        Thinobius fimbriatus LeConte, 1877 - Downie and Arnett 1996

References - Distribution

    Archangelsky M, Branham MA. 1998. Description of the preimaginal stages of Pyractomena borealis (Randall, 1838)(Coleoptera: Lampyridae) and notes on its biology. Proceedings of the Entomological Society of Washington 100(3):421-430).
    Askevold IS. 1991. Classification, reconstructed phylogeny, and geographic history of the new world members of Plateumaris Thomson, 1859 (Coleoptera: Chrysomelidae: Donaciinae). Memoirs of the Entomological Society of Canada 157:1-175.
    Atton FM. 1990. Gyrinus (Gyrinulus) cavatus sp. nov. from North America described and compared with Gyrinus (Gyrinulus) minutus Fabricius (Coleoptera: Gyrinidae). The Canadian Entomologist 122(4):651-657.
    Brigham WU. 1981. Ectopria leechi, a new false water penny from the United States (Coleooptera: Eubriidae). Pan-Pacific Entomologist 57:313-320.
    Brown HP. 1972. Aquatic dyropoid beetles (Coleoptera) of the United States. Biota of Freshwater Ecosystems Identification Manual No. 6. United States Environmental Protection Agency Project # 18050, Contract # 14-12-894, April 1972. x + 82 pp.
    Brown HP. 1987. Biology of riffle beetles. Annual Review of Entomology 32: 253-273.
    Brown HP, Murvosh CM. 1974. A revision of the genus Psephenus (water-penny beetles) of the United States and Canada (Coleoptera, Dryopoidea, Psephenidae). Transactions of the American Entomological Society 100:289-340.
    Cheary BS. 1971. The biology, ecology and systematics of the genus Laccobius (Laccobius) (Coleoptera: Hydrophilidae) of the new world. Privately printed. Riverside, CA. xv + 177 pp.
    Creed RP. 1998. A biogeographic perspective on Eurasian watermilfoil declines: additional evidence for the role of herbivorous weevils in promoting declines?. Journal of Aquatic Plant Management 36: 16-22.
    Downie NM, Arnett RH. 1996. The beetles of Northeastern North America. Volume I. Introduction; Suborders Archostemata, Adephaga, and Polyphaga, thru Superfamily Cantharoidea. The Sandhill Crane Press: Gainesville, Florida. xiv + 880 pp.
    Funk DH, Fenstermacher H. 2002. Description of the aquatic larva and pupa of Paralichas trivittis (Coleoptera: Ptilodactylidae), with notes on its biology. Entomological News 113(5): 344-354.
    Hardy AR, Bheary BS, Malcolm SE. 1981. A clarification of the authorship of some names in Laccobius Erichson (Coleoptera: Hydrophilidae). Pan-Pacific Entomologist 57(1):303-305.
    Hilsenhoff WL. 1992. Dytiscidae and Noteridae of Wisconsin (Coleoptera). I. Introduction, key to genera of adults, and distribution, habitat, life cycle, and identification of species of Abagetinae, Laccophilinae and Noteridae. The Great Lakes Entomologist 25(2): 57-69.
    Hilsenhoff WL. 1995b. Aquatic Hydrophilidae and Hydraenidae of Wisconsin (Coleoptera). I. Introduction, key to genera, and distribution, habitat, life cycle and identification of species of Helophorus Fabricius, Hydrochus Leach, and Berosus Leach (Hydrophilidae), and Hydraenidae. The Great Lakes Entomologist 28(1):25-53.
    Hilsenhoff WL. 1995c. Aquatic Hydrophilidae and Hydraenidae of Wisconsin (Coleoptera). II. Distribution, habitat, life cycle and identification of species of Hydrobiini and Hydrophilini (Hydrophilidae: Hydrophilinae). The Great Lakes Entomologist 28(2):97-126.
    Jolivet P, Hawkeswood TJ. 1995. Host-plants of Chrysomelidae of the World: An Essay about the Relationships Between the Leaf-beetles and their Food-plants. Backhuys Publishers: Leiden, Netherlands. 281 pp. + xiii.
    Jolivet P., Verma KK. 2002. Biology of leaf beetles. Intercept Limited: Andover, UK. 332 pp. + xiv
    Kölsch G., Pedersen BV. 2008. Molecular phylogeny of reed beetles (Col., Chrysomelidae, Donaciinae): The signature of ecological specialization and geographical isolation. Molecular Phylogenetics and Evolution 48: 936–952.
    Lloyd JE. 1999. On research and entomological education III: firefly brachyptery and wing "polymorphism" at Pitkin Marsh and watery retreats near summer camps (Coleoptera: Lampyridae; Pyropyga. Florida Entomologist 82(2):165-179
    Luk SPL, Marshall SA, Branham MA. 2011. The fireflies of Ontario (Coleoptera: Lampyridae). Canadian Journal of Arthropod Identification 16: 1-105.
    Merritt RW, Benbow ME, Hudson PL. 2002. Wetland macroinvertebrates of Prentiss Bay, Lake Huron, Michigan: diversity and function group comparison. The Great Lakes Entomologist 35(2): 149-160.
    Musgrave PN. 1935. A synopsis of the genus Helichus Erichson in the United States and Canada, with description of a new species (Coleoptera: Dryopidae). Proceedings of the Entomological Society of Washington 37(7):137-144, 1 pl.
    Oygur S, Wolfe GW. 1991. Classification, distribution, and phylogeny of North American (North of Mexico) species of Gyrinus Müller (Coleoptera: Gyrinidae). Bulletin of the American Museum of Natural History 207:1-97.
    Perkin PD. 1980. Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae 16(1-2):3-554.
    Ryker LC. 1972. Acoustic behavior of four sympatric species of water scavenger beetles (Coleoptera, Hydrophilidae, Tropisternus). Occasional Papers of the Museum of Zoology, University of Michigan 666:1-19.
    Smetana A. 1974. Revision of the genus Cymbiodyta Bed. (Coleoptera: Hydrophilidae). Memoirs of the Entomological Society of Canada 93: 1-113.
    Smetana A. 1980. Revision of the genus Hydrochara Berth. (Coleoptera: Hydrophilidae). Memoirs of the Entomological Society of Canada 111:1-100.
    Smetana A. 1988. Review of the family Hydrophilidae of Canada and Alaska (Coleoptera). Memoirs of the Entomological Society of Canada 142. 316 p.
    Tanner VM. 1943. A study of the subtribe Hydronomi with a description of new species, (Curculionidae), study No. VI. The Great Basin Naturalist 3(1-2):1-38.
    Waltz RD. 2002. Reports of rarely collected or previously unreported Indiana Coleoptera (Insecta): Cerambycidae, Cerophytidae, Hydraenidae, Hydrophilidae, Nitidulidae, Scarabaeidae, Staphylinidae, Trogossitidae. Proceedings of the Indiana Academy of Science 111(2): 177-181.
    Willson RB. 1970. The Hydrophilidae of Michigan with keys to species of the Great Lakes region. MS Thesis, Michigan State University, East Lansing, Michigan. vi + 100 p.
    Yanoviak SP, McCafferty WP. 1996. Comparison of macroinvertebrate assemblages inhabiting pristine streams in the Huron Mountains of Michigan, USA. Hydrobiologia 330: 195-211.
    Young FN. 1981. Predaceous water beetles of the genus Desmopachria: the convexa-grana group. Occasional Papers of the Florida State Collection of Arthropods 2: 1-11.
    Young DK, Stribling JB. 1990. Systematics of the North American Cyphon collaris species complex with the description of a new species (Coleoptera: Scirtidae). Proceedings of the Entomological Society of Washington 92(2): 194-204.

Page edits:

    August 08, 2016
    • Added new records of Plateumaris (Chrysomelidae, Donaciinae) from publication records in Askevold 1991
    August 04, 2016
    • Added new records of Chrysomelidae from publication records in Kölsch & Pedersen 2008
    January 16, 2013
    • Added Phaenonotum exstriatum to list; Smetana (1988) indicates a semi-aquatic ecology, found along water edges; Willston (1970) found it Sphagnum bog mat
    • Added species of Cercyon that Smetana (1988) considers associated with aquatic habitats (peri- to semi-aquatic)
    November 21, 2012
    • Beginning to keep track of changes to the web site pages
    • Removed individual pages for families into one long list, families accessed from left menu via anchor redirects within page.

Page created: November 01, 2002; Last update July 21, 2017 (EB)