Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

Semiaquatic Staphylinidae (Rove Beetles) of Michigan

This large family of slender beetles are characteristic in having short, truncated elytra that leaves several abdominal segments dorsally visible. The great majority of species are terrestrial, but a few are associated with aquatic environments (peri- and semiaquatic), principally with shore-line environments and the surface of littoral habitats. Of the groups known to be associated with aquatic habitats, there are 70 species in 6 genera found in Michigan, with another 9 species that may be found based on regional records. Psephidonus is common under stones at or near the edge of running water. Carpelimus and Thinobius is generally found along littoral areas of lakes and rivers among leaf litter. Not all species of Bledius and Stenus are semiaquatic/periaquatic, and provisionally terrestrial taxa are included in the list based on citations contained in the literature listed; museum and other collection specimens have have only begun to be examined.

Genera are donoted in bold green; species denoted in bold blue have been recorded in Michigan; other species denoted in bold are likely to occur based on their known distribution. You can use the Find function in your web browser to locate families, genera, and species. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated.

Bledius Leach, 1819: 174 (Subfamily Oxytelinae)

Bledius analis LeConte, 1863: 52 - Herman 1972: 231
Bledius annularis LeConte, 1863: 53 - this is a large species complex (incl. B. annularis LeConte and B. sinuatus LeConte 1877: 228) widely distributed in NA - see Herman 1983: 94-95 for discussion
Bledius assimilis Casey, 1889: 55 - Herman 1972: 243
Bledius confusus LeConte, 1877: 228 - Downie and Arnett 1996: 455, WI (Herman 1983: 124)
Bledius emarginatus (Say, 1834: 461 as Oxytelus) - Herman 1983: 15
(syn.) Bledius troglodytes Erichson, 1840: 774
Bledius fumatus LeConte, 1863: 52 - Herman 1972: 221
Bledius gravidus Casey, 1889: 54 - Herman 1972: 217
Bledius ineptus Casey, 1889: 48 - Herman 1976: 124
Bledius nitidicollis LeConte, 1863: 52 - IL, OH, and northwest, expect in sw LP (Herman 1972: 225)
Bledius omega Herman, 1983: 45 - Herman 1983: 47
Bledius ruficornis LeConte, 1863: 53 - Herman 1983: 84
Bledius semiferrugineus LeConte, 1863: 52 - UMMZ Insect Collection; Herman 1972: 205
(syn.) Bledius canaliculatus Notman 1920: 697
Bledius suturalis LeConte, 1863: 54 - ON, widespread, expect in UP (Herman 1983: 54)
(syn.) Bledius luteipennis LeConte, 1877: 227
(syn.) Bledius pleuralis LeConte, 1877: 229
(syn.) Bledius medialis Fall, 1910: 113
(syn.) Bledius oregonensis Hatch, 1957: 103
Bledius tarandus Herman, 1970: 379 - Herman 1983: 36
(syn.) Bledius divisus LeConte, 1863: 53
Bledius tau LeConte, 1877: 226 - Herman 1983: 45
Bledius trugidus Casey, 1889: 52 - Herman 1983: 71
(syn.) Bledius borealis Blatchley, 1910: 465
(syn.) Bledius bowronensis Hatch, 1957: 105
Bledius viriosus Herman, 1983: 73 - ON, WI (Herman 1983: 75)

Carpelimus Leach, 1819: 174 (Subfamily Oxytelinae)

Carpelimus delicatus (Casey, 1889: 365 as Trogophloeus) - Casey 1889: 365, (Detroit, Wayne Co.)
Carpelimus facetus (Casey, 1889: 378 as Trogophloeus) - Casey 1889: 375 as T. fallax, Lake Superior)
(syn.) Trogophloeus fallax Casey, 1889: 375
Carpelimus morio (Erichson, 1840: 805 as Trogophloeus) - Downie and Arnett 1996: 444
Carpelimus quadripunctatus (Say, 1831: 46 [= 1834: 459] as Stenus) - IN, OH (Blatchley, 1910: 471)

Psephidonus Gistel, 1856: 29 (=Geodromicus Redtenbacher) (Subfamily Omaliinae)

Psephidonus brunneous (Say, 1823: 158 as Anthophagus) - UMMZ Insect Division
(syn.) Anthophagus caesus Erichson, 1840: 853
Psephidonus ovipennis (LeConte, 1878: 452 as Geodromicus) - widespread, disjunct (New York, Utah)
Psephidonus strictus (Fauvel, 1889: 126 as Geodromicus) - Casey 1894: 408, Herman 2001a: 312


Stenus Latreille, 1797: 77 (Subfamily Steninae)

Stenus (Hypostenus) advena (Casey, 1884: 158 as Areus) - Casey 1884, holotype Whitefish Pt. and Marquette, Lake Superior; Puthz 1972a: 14; Herman 2001b: 2045
Stenus (Stenus) angustus Casey, 1884: 40 - Casey 1884: 65 (as S. placidus), Casey 1884: 100 (as S. tumicollis), Puthz 1972: 12; Herman 2001b: 2058
(syn.) Stenus indigens Casey, 1884: 55
(syn.) Stenus pettiti Casey, 1884: 42
(syn.) Stenus placidus Casey, 1884: 65
(syn.) Stenus mendax Casey, 1884: 66
(syn.) Stenus tumicollis Casey, 1884: 100
Stenus (Stenus) annularis Erichson, 1840: 735 - Casey 1884: 168 (as Areus annularis), Puhtz 1972: 13, Puhtz 1974: 11
Stenus (Hypostenus) arculus Erichson, 1840: 744 - Casey 1884: 188 (as Areus arculus)
Stenus (Nestus) brivioi Puthz, 1972a: 12-13 - Puthz 1972a: 13 (Maryglade College, E. of Memphis, Macomb County)
Stenus (Hypostenus) callosus Erichson, 1840: 737 - Puthz 1972a: 14
Stenus (Nestus) canaliculatus Gyllenhal, 1827: 501- Casey 1884: 115
(syn.) Stenus affinis Stephens, 1833: 298
(syn.) Stenus foveifrons Rey, 1884: 272
(syn.) Stenus cordatifollis Leinberg, 1900: 187
Stenus (Hypostenus) capucinus Boheman, 1858: 34 - Fauvel 1901: 76, Downie and Arnett 1996: 494
(syn.) Stenus micans Sharp, 1886: 656
(syn.) Stenus sandersoni L. Benick, 1941: 222
Stenus (Stenus) carinicollis Casey, 1884: 101 - Casey 1884: 101 (Marquette, holotype)
Stenus (Nestus) caseyi Puthz, 1972b: 172 - Puthz 1972b: 172, Puthz 1975: 125
Stenus (Stenus) colon Say, 1831: 44 [1834: 458] - Casey 1884: 14
(syn.) Stenus carolinus Bernhauer, 1917: 1

Stenus (Nestus) colonus Erichson, 1840 - Puthz 1972a: 12 (Lapeer Co, 1 mi. S of Lum), Puthz 1974: 9
(syn.) Stenus arizonae Casey, 1884: 93
(syn.) Stenus militaris Casey, 1884: 69
(syn.) Stenus pacificus Casey, 1884: 91
(syn.) Stenus suspectus Blatchley, 1910
(syn.) Stenus obstrusus Casey, 1884: 92
(syn.) Stenus integer Casey, 1884: 94 (syn.) Stenus floridanus Casey, 1884: 95
(syn.) Stenus championi Sharp, 1886: 642
(syn.) Stenus inermis Sharp, 1886: 642
(syn.) Stenus suspectus Blatchley, 1910: 408
(syn.) Stenus interpres L. Benick, 1926: 264
(syn.) Stenus weiseri Bernhauer, 1927: 234
(syn.) Stenus chapini Blackwelder, 1943: 211

Stenus (Stenus) comma LeConte, 1863: 50 - Puhtz 1972: 11
(syn.) Stenus bipustulatus Stephens, 1833: 303
(syn.) Stenus bipunctatus Erichson, 1839: 530
(syn.) Stenus punctiger Casey, 1884: 13
(syn.) Stenus aeneiceps Rey, 1884: 192
(syn.) Stenus ananias Bondroit, 1912: 263
(syn.) Stenus peroculatus Puthz, 1986: 419
Stenus (Hemistenus) croceatus (Casey, 1884: 158 as Areus) - Casey 1884: 158 (Detroit, Marquette), Sanderson 1947: 427, Puthz 1972a: 13 (Macomb Co., 1 mi. E. of Mephis; ESGR, Livingston Co., Ann Arbor, Washtenaw Co.)
Stenus (Stenus) curtus Casey, 1884: 135 - Casey 1884: 135 (Whitefish Pt., Chippewa Co.)
Stenus (Stenus) difficilis Casey, 1884: 41 - Casey 1884: 54, 56, 67 (as S. subtilis, S. subtilis, S. subtilis - Detroit, Wayne Co.), Puthz 1972a: 12 (1 mi. E of Memphis, Macomb Co.)
(syn., preocc.) Stenus tenuis Casey, 1884: 40
(syn.) Stenus pauper Casey, 1884: 53
(syn.) Stenus subtilis Casey, 1884: 54
(syn.) Stenus subtilis Casey, 1884: 58
(syn.) Stenus inornatus Casey, 1884: 64
(syn.) Stenus subtilis Casey, 1884: 67
(syn.) Stenus parallelus Casey, 1884: 121
Stenus (Stenus) dissentiens (Casey, 1884: 190 as Areus) - Casey 1884: 190 (Whitefish Pt., Chippewa Co.; also as A. simiolus), Sanderson 1957: 250
(syn.) Areus hubbardi Casey, 1884: 191
(syn.) Areus leviceps Casey, 1884: 193
(syn.) Areus politulus Casey, 1884: 131
(syn.) Areus simiolus Casey, 1884: 194

Stenus (Nestus) egenulus Puthz, 1988: 145 - Puthz 1988: 145
Stenus (Nestus) egenus Erichson, 1840: 698 - LeConte 1850: 220 (Lake Superior), Casey 1884: 105, Puthz 1972a: 12 (1 mi. E of Mephis, Macomb Co; Port Sanilac, Sanilac Co., Douglas Lake, Emmet Co., Ann Arbor, Washtenaw Co.)
Stenus (Stenus) erythropus Melsheimer, 1844: 40 - Casey 1884: 46, Puthz 1972a: 11 (ESGR, Livingston Co.)
(nom nud., preocc.) Laestris fuscipes Melsheimer, 1844: 40

Stenus (Stenus) femoratus Say, 1834: 45 [1843: 459] - Casey 1884: 33, Puthz 1972a: 12 (1 mi. S of Lum, Macomb Co.), Puthz 1974: 9
Stenus (Hypostenus) flavicornis Erichson, 1840 - Casey 1884: 167 (as Areus, Lake Superior), Puthz 1972a: 13 (Maryglade College, 1 mi. E of Memphis, Macomb Co., ESGR, Livingston Co., Harbor Beach, Huron Co., Douglas Lake, Emmet Co., Ann Arbor, Washtenaw Co.)
(preocc.) Stenus geniculatus Say, 1831: 44

Stenus (Stenus) fulvoguttatus Notman, 1920: 190 - Downie and Arnett 1996: 489
Stenus (Tesnus) gratiosus Casey, 1884: 133 - Casey 1884: 133 (Holotype, Detroit, Wayne Co.; also as S. hirsutus), Puthz 1972a (Maryglade College, 1 mi. E of Memphis, Macomb Co.)
(syn.) Stenus hirsutus Casey, 1884: 137
Stenus (Tesnus) immarginatus Mäklin, 1853: 192 - ON, widespread north, may expect in UP (Campbell and Davies 2013 PDF)
(syn.) Stenus monticola Casey, 1884: 134
(syn.) Stenus variabilis Puthz, 1968: 207
Stenus (Stenus) intrusus Casey, 1884: 44 - Casey 1884: 44 (Whitefish Pt., Chippewa Co.; Escanba, Delta Co.)
(syn.) Stenus simpliciventris Poppius, 1909: 13
(syn.) Stenus guranicus Ryvkin, 1987: 149
Stenus (Hypostenus) insperatus Puthz, 1972: 14 - Puhtz 1972 (Grand Ledge, Eaton Co.)
Stenus (Stenus) juno Paykull, 1789: 33 - Casey 1884: 24, Puthz: 12 (1 mi. E. of Mephis, Macomb Co.; Port Salinac, Salinac Co.; Harbor Beach, Huron Co.), very widespread Holarctic species (Puthz 1972a, Herman 2001b: 2240-2242)
(syn.) Stenus clavicornis Fabricius, 1777: 242
(syn.) Stenus buphthalmus Latreille, 1804: 353
(syn.) Stenus maurus Mannerheim, 1830: 41
(syn.) Stenus lineatulus Stephens, 1833: 296
(syn.) Stenus calabrus Fiori, 1894: 95
Stenus (Stenus) laccophilus Casey, 1884 - UMMZ insect collection, widespread north (Campbell and Davies 2013 PDF), Herman 2001b: 2251
(syn.) Stenus verticosus Casey, 1884: 25
(syn.) Stenus rugifer Casey, 1884: 47
(syn.) Stenus anastomozans Casey, 1884: 48
(syn.) Stenus edax Notman, 1920: 191
Stenus (Nestus) mammops Casey, 1884: 97 - Casey 1884: 97 (Detroit, Wayne Co.; Marquette, Marquette Co.; Isle Royale, Keweenaw Co.), Puthz 1972a: 13 (ESGR, Livingston Co.)
(syn.) Stenus bulbicollis L. Benick, 1921b: 151
(syn.) Stenus rulomus Blackwelder, 1943: 212
Stenus (Nestus) morio Gravenhorst, 1806: 230 - Casey 1884: 125 (as S. haplus, Detroit, Wayne Co.), Puthz 1972a: 12 (Macomb Co.)
(syn.) Stenus aequalis Mulsant and Rey, 1861: 154 [= 1862: 138]
(syn.) Stenus inaequalis Mulsant and Rey, 1861: 156 [= 1862: 140]
(syn.) Stenus luculentus Casey, 1884: 125
(syn.) Stenus haplus Casey, 1884: 125
(syn.) Stenus subgriseus Casey, 1884: 127
(syn.) Stenus dives Casey, 1884: 127
(syn.) Stenus terricola Casey, 1884: 128
(syn.) Stenus indistinctus Casey, 1884: 130
(syn.) Stenus albipilus Rey, 1884: 275 [= 1884a: 123]
(syn.) Stenus arcuatus Rey, 1884: 282
Stenus (Nestus) nanus Stephens, 1833: 301 - Downie and Arnett 1996: 489
(syn.) Stenus declaratus Erichson, 1839: 557
Stenus (Stenus) neglectus Casey, 1884: 59 - Puthz 1972a: 12 (1 mi. E. of Memphis, Macomb Co.; Ann Arbor, Washtenaw Co.)
Stenus (Stenus) pauperculus Casey, 1884: 124 - Casey 1884: 124 (probably from Michigan, also as S. brumalis from Detroit, Wayne Co.; Ann Arbor, Washtenaw Co.), Puthz, 1971: 244
(syn.) Stenus brumalis Casey, 1884: 131
Stenus (Hemistenus) pubescens Stephens, 1833: 297 - Casey 1884 (as S. fraternus, Marquette, Marquette Co.), Puthz, 1972c: 84, 88
(syn.) Stenus laevior Stephens, 1833: 297
(syn.) Stenus laevis Stephens, 1833: 297
(syn.) Stenus subimpressus Erichson, 1839: 561
(syn.) Stenus fraternus Casey, 1884: 155
(syn.) Stenus sandersonianus Puthz, 1972c: 82
Stenus (Stenus) pudicus Casey, 1884: 117 - Casey 1884: 117 (Whitefish Pt., Chippewa Co.), Casey 1884: 118, 123 as S. minor, S. juvencus - Detroit, Wayne Co.)
(syn.) Stenus minor Casey, 1884: 118
(syn.) Stenus juvencus Casey, 1884: 123
(syn.) Stenus enodis Casey, 1884: 126
(syn.) Stenus sibyllinus L. Benick, 1921a: 117
Stenus (Stenus) pumilio Erichson, 1839: 558 - Casey 1884: 84 (as S. atomarius, Detroit, Wayne Co.)
(syn.) Stenus atomarius Casey, 1884: 84
Stenus (Hypostenus) punctatus Erichson, 1840: 744 - Puthz 1974: 11
(syn.) Stenus obsoletus Casey, 1884: 181
(syn.) Stenus artus Casey, 1884: 182
(syn.) Stenus dilutus Casey, 1884: 184
Stenus (Nestus) pusillimus Puthz, 1971: 250, 254 - Puthz, 1971: 250
Stenus (Hypostenus) reconditus (Casey, 1884: 173 as Areus) - Puthz, 1972d: 317
(syn.) Stenus propinquus Casey, 1884: 174
(syn.) Stenus canadensis Casey, 1884: 175
Stenus (Hemistenus) retrusus (Casey, 1884: 154 as Areus) - western, but record from WI noted by Puthz, 1972c: 87
Stenus (Stenus) rossi Sanderson, 1957: 238 - Sanderson 1957: 238 (Cheboygan Co., Oakland Co.)
Stenus (Stenus) scabiosus Casey, 1884: 60 - Casey 1884: 60 (Lake Superior, Marquette, Marquette Co.)
Stenus (Stenus) scrupeus Casey, 1884: 43 - Casey 1884: 43 (Lake Superior, Marquette, Marquette Co.)
Stenus (Stenus) semicolon LeConte, 1863a: 50 - Casey 1884: 17 (Lake Superior)
Stenus (Parastenus) sibiricus Sahlberg, 1880: 79 - Casey 1884: 17 (as S. confusus- Whitefish Pt., Lake Superior, Chippewa Co.)

(syn.) Stenus confusus Casey, 1884: 63
(syn.) Stenus montanus Casey, 1884: 74
(syn.) Stenus deductor Casey, 1884: 204
(syn.) Stenus exasperatus L. Benick, 1925: 73
(syn.) Stenus wenzeli Puthz, 1967: 239
Stenus (Stenus) strangulatus Casey, 1884: 39 - Casey 1884: 39 (Detroit, Wayne Co.)
Stenus (Stenus) stygicus Say, 1831: 45 [= 1834: 458] - Casey 1884: 104 (Lake Superior, Marquette, Marquette Co.), Puthz 1972a (1 mi. E of Memphis, Macomb Co.; Douglas Lake, Emmet Co.)
Stenus (Hypostenus) varipes (Casey, 1884: 187 as Areus) - Casey 1884: 187, Detroit, Wayne Co.
Stenus (Stenus) vicinus Casey, 1884: 57 - Puthz, 1972a: 12 (1 mi E. of Memphis, Macomb Co.)
Stenus (Nestus) vinnulus Casey, 1884: 112 - Casey 1884: 112 (White Fish Point, Chippewa Co.; Isle Royale, Keweenaw Co., Lake Superior; Marquette, Marquette Co.)
(syn.) Stenus confusoides Renkonen, 1935: 27

Thinobius Kiesenwetter, 1844: 355 (Subfamily Oxytelinae)

Thinobius brachypterus (LeConte, 1877: 240 as Carpelimus) - LeConte 1877: 240 (holotype, Detroit), Downie and Arnett 1996: 456
(syn.) Thinobius brevicollis Mulsant and Rey, 1870
Thinobius fimbriatus LeConte, 1877: 240 - LeConte 1877: 240-241 (holotype, Detroit), Fauvel, 1878: 263, Lake Superior), Casey 1889b: 84

References - Distribution

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Page edits:

July 13, 2020
• Updated species list
• Removed Carpelimus bilineatus, Stenus alacer, S. pallipennis, S. parallelopipedus and Bledius rotundicollis, not likely to occur in MI; specimen in the UMMZ of Bledius armatus Say, which is replaced by B. bellicus Blackwelder, which is found in the central USA (see Herman 1976: 133) and not likely to occur in Michigan
• Stenus annularis is a species complex (see Herman 1983: 122 for a list) - B. annularis LeConte and B. sinuatus LeConte likely to occur in Michigan
November 21, 2012
• Beginning to keep track of changes to the web site pages
• Removed individual pages for families into one long list, families accessed from left menu via anchor redirects within page.

Page created: November 01, 2002; Page last updated: July 15, 2020 (EB)