Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

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Dytiscidae (Predaceous Diving Beetles) of Michigan

Introduction

The Nearctic region has been relatively well surveyed, and most of this information has been recently included in a new publication by Larson et al. (2000). This list and nomenclature is largely based on that work as well as provisional records in the UMMZ and the Smithsonian (NMNH web page database). Genera are donoted in bold green; species denoted in bold blue have been recorded in Michigan; other species denoted in bold are likely to occur based on their known distribution. You can use the Find function in your web browser to locate families, genera, and species. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated.

Photo of Bidessonotus sp. adult from Honduras, Copyright © 2006 Ethan Bright
Bidessontus sp. from Honduras
    Acilius Leach, 1817: 69, 72 (Subfamily Dytiscinae; Tribe Aciliini)
      Acilius athabascae Larson, 1975: 410-412 - MN, boreal, may occur in Isle Royale NP, nw UP (Larson et al. 2000: 813-814)
      Acilius fraternus (Harris, 1828a: 156 as Dytiscus) - Hilsenhoff 1975: 273; see also Larson et al. 2000: 812
      Acilius mediatus (Say, 1823b: 93 as Dytiscus) - Hilsenhoff 1975: 273; see also Larson et al. 2000: 807-808
      Acilius semisulcatus Aubé, 1838b: 132 - Hilsenhoff 1975: 273-274; Larson et al. 2000: 808-810
      Acilius sylvanus Hilsenhoff, 1975: 271 - Hilsenhoff 1975: 271-273; Larson et al. 2000: 811-812

    Agabetes Crotch, 1873: 401 (Subfamily Agabetinae)
      Agabetes acuductus (Harris, 1828b: 164 as Colyembetes) - IN, ON, WI, expect in LP (Hilsenhoff 1992: 64-65; Larson et al. 2000: 56-58)
        (syn.) Agabus regulosis Aubé, 1838b: 362-363

    Agabus Leach, 1817: 68 (Subfamily Colymbetinae, Tribe Agabini)
      Agabus aeruginosus Aubé, 1838a: 298 - Larson et al. 2000: 596-597
      Agabus ambiguus (Say, 1823b: 96 as Colymbetes.) - Larson 1989: 879-883; Larson et al. 2000: 625-626
        (syn.) Agabus reticulatus Aubé, 1838b: 355
        (syn.) Agabus fimbriatus LeConte, 1850: 214
      Agabus antennatus Leech, 1939: 217 - MN, ON, WI, widespread north (Hilsenhoff 1993c: 181; Larson et al. 2000: 580-582)
        (syn.) Agabus clavatus LeConte, 1859a: 4
      Agabus anthracinus Mannerheim, 1852: 304 - Larson 1989: 896-898; Larson et al. 2000: 635-637
        (syn.) Agabus scapularis Mannerheim, 1852: 303
      Agabus bicolor (Kirby, 1837: 70 as Colymbetes) - Larson 1991: 1269-1271, 1314; Larson et al. 2000: 679-681
        (syn.) Agabus mutus Sharp, 1882: 513
      Agabus bifarius (Kirby, 1837: 71 as Colymbetes) - Larson et al. 2000: 589-591
      Agabus canadensis Fall, 1922: 27 - MN, ON, WI, expect in UP (Larson 1991: 1264-1266, 1313; Hilsenhoff 1993c: 183; Larson et al. 2000: 675-676)
      Agabus confinis (Gyllenhal, 1808: 511 as Dytiscus) - Larson 1991: 1287-1289, 1311; Larson et al. 2000: 692-694
        (nom.nud.) Agabus ovoideus LeConte, 1863: 17
        (syn.) Gaurodytes ovoideus Crotch, 1873: 418
        (syn.) Gaurodytes longulus LeConte, 1878: 596
      Agabus confusus (Blatchley, 1910: 229 as Rhantus) - WI (MI? found in south central WI: Hilsenhoff 1993c: 183)
        (syn.) Agabus ampulus Fall, 1922: 10, 12
      Agabus discolor (Harris, 1828b: 164 as Colymbetes) - Larson 1991: 1272-1275, 1315; Larson et al. 2000: 683-684
      Agabus disintegratus (Crotch, 1873: 416 as Gaurodytes) - UMMZ (see also Hilsenhoff 1993c: 184; Larson et al. 2000: 600-602)
      Agabus erichsoni Gemminger & Harold, 1868: 454 - Larson et al. 2000: 506-508
        (syn.) Agabus nigroaeneus Erichson 1837: 157, not Marsham 1802
      Agabus falli (Zimmermann, 1934: 186 as Gaurodytes) - IN, MN, ON, WI (Hilsenhoff 1993c: 185; Larson et al. 2000: 616-617)
        (syn.) Agabus semipunctatus Sharp, 1882: 507, not Kirby, 1837
        (syn.) Agabus sharpi Fall, 1922: 19
      Agabus flavovittatus Larson and Wolfe, 1998: 44 - IL, IN, OH, ON (expect in s. MI; see Larson et al. 2000: 542)
      Agabus fuscipennis (Paykull, 1798: 209 as Dytiscus) - ON (boreal, may occur in UP) (Hilsenhoff 1993c: 185; Larson 1994: 155-156, 180; Larson et al. 2000: 6583-654)
        (syn.) Agabus ontarionis Fall, 1922: 30
      Agabus gagates Aubé, 1838b: 306 - record at NMNH (web page database); see also Larson et al. 2000: 508-509
      Agabus immaturus Larson, 1991: 1263 - Larson 1991: 1263-1264, 1313; Larson et al. 2000: 673-675
      Agabus inscriptus (Crotch, 1873: 422 as Gaurodytes) - Larson 1991: 1281-1284, 1316; Larson et al. 2000: 688-689
        (syn.) Agabus smithi Brown, 1930a: 88
      Agabus larsoni Fery and Nilsson, 1993: 98 - ON (Larson et al. 2000: 509-510)
      Agabus leptapsis (LeConte, 1878: 596 as Gaurodytes) - NMNH (web page database); see also Larson 1989: 892-894; Larson et al. 2000: 632-634
      Agabus obtusatus (Say, 1823b: 99-100 as Colymbetes) - IN, OH, ON (Hilsenhoff 1993c: 187; Larson et al. 2000: 585-586)
      Agabus opacus Aubé, 1837: 173 - UMMZ, NMNH (web page database - as A. pseudoconfertus Wallis); Larson 1996: 631-633, 664; Larson et al. 2000: 521-523
        (syn.) Agabus sharpi Jacobson 1908 (replacement for A. sibericus Sharp 1882, not J. Shalberg 1880)
        (syn.) Gaurodytes mimmi Shalberg, 1875: 182
        (syn.) Agabus sibericus Sharp, 1882: 519
        (syn.) Agabus gelidus Fall, 1926: 142
        (syn.) Agabus pseudoconfertus Wallis, 1926: 90
        (syn.) Agabus sachalinensis Kamiya, 1938: 37
      Agabus phaeopterus (Kirby, 1837: 70 as Colymbetes) - Larson 1991: 1260-1262, 1312; Larson et al. 2000: 671-673
      Agabus punctatus Melsheimer, 1844: 27-28 - Larson et al. 2000: 594-595
      Agabus punctulatus Aubé, 1838b: 332 - IL, IN, MN, ON, WI (Hilsenhoff 1993c: 188; Larson et al. 2000: 609-610)
        (syn.) Gaurodytes aeneolus Crotch, 1873: 421
      Agabus semipunctatus (Kirby, 1837: 69 as Colymbetes) - Larson et al. 2000: 605-606
        (syn.) Agabus stridulator Sharp, 1882: 509
        (syn.) Agabus stridulatus Zimmermann, 1920: 173
      Agabus semivittatus LeConte, 1852: 204 - UMMZ
      Agabus seriatus (Say, 1823b: 97 as Colymbetes) - LeConte 1850: 213 (as A. parallelus); Larson et al. 2000: 559-561
        (syn.) Agabus striatus Aubé, 1838a: 305
        (syn.) Agabus arctus Melsheimer, 1844: 27
        (syn.) Agabus parallelus LeConte, 1850: 213
        (syn.) Gaurodytes intersectus Crotch, 1873: 419
      Agabus subfuscatus Sharp, 1882: 514 - Larson 1991: 1271-1272, 1317; Larson et al. 2000: 681-683
      Agabus thomsoni (Sahlberg, 1871: 407 as Gaurodytes) - Larson 1991: 1255-1257, 1309; Larson et al. 2000: 666-668
        (syn.) Agabus congener auctorum, not Thunberg 1794: 75
      Agabus wasastjernae (Sahlberg, 1824: 167 as Dytiscus) - Larson 1996: 629-631, 664; Larson et al. 2000: 519-521
        (syn.) Agabus kenaiensis Fall, 1926: 141
        (syn.) Agabus palustris Wallis, 1926: 92

    Bidessonotus Régimbart, 1895: 331 (Subfamily Hydroporinae, Tribe Bidessini)
      Bidessonotus inconspicuus (LeConte, 1855: 290 as Hydroporus) - UMMZ; see also Larson et al. 2000: 105-106
      Bidessonotus pulicarius (Aubé, 1838b: 494 as Hydroporus) - IN, ON (Larson et al. 2000: 107)

    Celina Aubé, 1837: 218 (Subfamily Hydroporinae, Tribe Methlini)
      Celina hubbelli Young, 1979: 826 - Young 1979: 826 (Lenawee Co.); Larson et al. 2000: 90-91

    Colymbetes Clairville, 1806: 188 (Subfamily Colymbetinae, Tribe Colymbetini)
      Colymbetes exaratus LeConte, 1862: 522 - IL, MN, WI, may occur in the UP (Zimmerman 1981: 13-16 (as C. e. exaratus), Larson et al. 2000: 770)
        (syn.) Colymbetes sculptilis exaratus, Young & Severin, 1956: 79
        (syn.) Colymbetes sculptilis (in part), Larson, 1975: 389, nec LeConte
        (syn.) Colymbetes rugipennis Sharp, 1882: 682
      Colymbetes paykulli Erichson, 1837: 149 - Zimmerman 1981: 40-41; Larson et al. 2000: 765-766
        (syn.) Colymbetes longulus LeConte, 1862: 522
      Colymbetes sculptilis Harris, 1829: 8 - Zimmerman 1981: 11, 18-20, 22-27; Larson et al. 2000: 767-769

    Copelatus Erichson, 1832: 18, 38 (Subfamily Copelatinae)
      Copelatus chevrolati Aubé, 1838b: 389 - Larson et al. 2000: 48, 50-51
        (syn.) Copelatus renovatus Guignot, 1952: 170
      Copelatus glyphicus (Say, 1823b: 99 as Colymbetes) - Larson et al. 2000: 51-52

    Coptotomus Say, 1834: 443 (Subfamily Colymbetinae, Tribe Coptotomini)
      Coptotomus longulus LeConte, 1852: 205 - Hilsenhoff 1980: 465-466 (as C. lenticus); Larson et al. 2000: 734-736
        (syn.) Coptotomus lenticus Hilsenhoff, 1980: 465
      Coptotomus loticus Hilsenhoff, 1980: 467 - IL, IN, ON, WI (Hilsenhoff 1980: 467-468; Larson et al. 2000: 736-737)

    Cybister Curtis, 1827: 151 (Subfamily Dytiscinae, Tribe Cybistrini)
      Cybister fimbriolatus (Say, 1823b: 91 as Dytiscus) - UMMZ, NMNH (web page database); see also Larson et al. 2000: 832-833

    Desmopachria Babington, 1841: 16 (Subfamily Hydroporinae, Tribe Hyphydrini)
      Desmopachria (Desmopachria) convexa (Aubé, 1838b: 479 as Hydroporus) - UMMZ, NMNH (web page database); Larson et al. 2000: 99-100; see also Young 1981: 3; Hilsenhoff 1994: 277

    Dytiscus Linnaeus, 1758: 411 (Subfamily Dytiscinae, Tribe Dytiscini)
      Dytiscus alaskanus Balfour-Browne, 1944: 77 - MN, ON (near Sault Ste. Marie), WI, expect in UP and n. LP (Roughley 1990: 481-483, 500; Hilsenhoff 1993a: 41; Larson et al. 2000: 792-793)
        (syn.) Dytiscus parvulus Motschulsky, 1852: 77
        (misid.) Dytiscus ooligbuckii J. Balfour-Browne 1948: 161, Hatch 1953: 238, Kirby 1837: 74
      Dytiscus carolinus Aubé, 1838b: 120 - Roughley 1990: 446-447, 453
        (syn.) Dytiscus fasciventris auctorim, nec Say, 1824: 270
        (syn.) ?Dytiscus fasciventris, Folkerts, 1978: 346
        (syn.) Dytiscus sp., Young, 1954: 26
      Dytiscus circumcinctus Ahrens, 1811: 55 - Roughley 1990: 477-481, 498; Larson et al. 2000: 791-792
        (syn.) Dytiscus flavocinctus Hummel, 1823: 17
        (syn.) Dytiscus angustatus Stephens, 1828: 88
        (syn.) Dytiscus dubius Gyllenhal, 1827: 372
        (syn.) Dytiscus ooligbuckii Kirby, 1837: 74
        (syn.) Dytiscus circumscriptus Boisduval & Lacordaire, 1835: 300
        (syn.) Dytiscus anxius Mannerheim, 1843: 218
        (syn.) Dytiscus fusco-striatus Motschulsky, 1853: 167
        (syn.) Dytiscus confusus Motschulsky, 1860: 101
        (misid.) Dytiscus albinicus auctorum, nec Motschulsky, 1859: 166
        (misid.) Dytiscus dauricus Anderson 1962: 73, nec Gebler, 1832: 39
      Dytiscus cordieri Aubé, 1838b: 108 - Roughley 1990: 456-458, 463; Larson et al. 2000: 789-790
        (syn.) Dytiscus sublimbatus LeConte, 1857: 34
        (misid.) Dytiscus marginalis auctorum, nec Linnaeus 1758: 411
      Dytiscus dauricus Gebler, 1832: 39 - Roughley 1990: 483-486, 501; Larson et al. 2000: 793-795
        (syn.) Dytiscus confluens Say, 1834: 440 (type lost)
        (syn.) Dytiscus franklinii Kirby, 1837: 77
        (nom.nud.) Dytiscus frontalis Motschulsky, 1859: 489
        (syn.) Dytiscus vexatus Sharp, 1882: 643
        (syn.) Dytiscus dauricus var. obscurus Gschwendtner, 1922: 93
        (syn.) Dytiscus amurensis J. Balfour-Browne 1944: 356 (new name for D. frontalis Motschulsky 1860 nee Marsham 1802)
      Dytiscus fasciventris Say, 1824: 270 - Roughley 1990: 447-449, 454; Larson et al. 2000: 786-788
      Dytiscus harrisii Kirby, 1837: 76 - Roughley 1990: 433-434, 440; Larson et al. 2000: 782-783
      Dytiscus hybridus Aubé, 1838b: 116 - Roughley 1990: 434-435, 441; Larson et al. 2000: 783-785
        (syn.) Leionotus compar Melsheimer, 1844: 26
      Dytiscus verticalis Say, 1823b: 92 - Roughley 1990: 431-433, 439; Larson et al. 2000: 780-782

    Graphoderus Dejean, 1833: 61 (Subfamily Dytiscinae, Tribe Aciliini)
      Graphoderus fasciocollis (Harris, 1828a: 156 as Dytiscus) - Larson et al. 2000: 821-822
      Graphoderus liberus (Say, 1825: 160 as Dytiscus) - Larson et al. 2000: 817-818
        (syn.) Dytiscus thoracicus Harris, 1828a: 156
        (syn.) Colymbetes rugicollis Kirby, 1837: 73
        (syn.) Hydaticus brunnipennis Aubé, 1838b: 203
      Graphoderus manitobensis Wallis, 1933: 276 - IL, WI, expect in western UP (Hilsenhoff 1993a: 47-48; Larson et al. 2000: 822-823)
      Graphoderus occidentalis Horn, 1883: 281 - NMNH (web page database); see also Larson et al. 2000: 820-821
      Graphoderus perplexus Sharp, 1882: 695 - Larson et al. 2000: 818-819
        (syn.) Graphoderus elatus Sharp, 1882: 695
        (misid.) Graphoderus cinereus Horn, 1883: 280 (not Linnaeus, 1758)
        (misid.) Graphoderus zonatus Zimmermann 1919: 231 (not Hoppe, 1795)
        (misid.) Graphoderus zonatus Hatch 1928 (not Hoppe, 1795)

    Heterosternuta Strand, 1935: 156, 161, 178 (Subfamily Hydroporinae, Tribe Hydroporini)
      Heterosternuta pulcher (LeConte, 1855: 298 as Hydroporus) - OH, ON, MN (Hilsenhoff 1995a: 4; Larson et al. 2000: 235-236)
      Heterosternuta wickhami (Zaitzev, 1908: 118 as Hydroporus) - UMMZ; see also Larson et al. 2000: 232-234

    Hydaticus Leach, 1817: 69 (Subfamily Dytiscinae, Tribe Hydaticini)
      Hydaticus aruspex Clark, 1864: 212 - Roughley and Pengelly 1981: 260-267, 300 (as H. modestus); Larson et al. 2000: 800-801
        (syn.) Hydaticus modestus Sharp, 1882: 650
      Hydaticus piceus LeConte, 1863: 23 - Roughley and Pengelly 1981: 267-268, 302; Larson et al. 2000: 802-803

    Hydrocolus (=Hydroporus oblitus-group Fall, 1923) Roughley & Larson, in Larson, Alarie & Roughley 2000: 407 (Subfamily Hydroporinae, Tribe Hydroporini)
      Hydrocolus paugus (Fall, 1923: 50 as Hydroporus) - Larson et al. 2000: 414-416
      Hydrocolus persimilis (Crotch, 1873: 395 as Hydroporus) - Larson et al. 2000: 416-417
        (syn.) Hydroporus aequus Fall, 1923: 53
        (syn.) Hydroporus acadianus J Balfour-Brown, 1948: 163
      Hydrocolus rybyae (Larson, 1975: 290 as Hydroporus) - ON, WI (boreal, expect in UP; Hilsenhoff 1995a: 16; Larson et al. 2000: 422-423)
      Hydrocolus stagnalis (Gemminger & Harold, 1868: 441 as Hydroporus (orig.) - Larson et al. 2000: 426-428
        (syn.) Hydroporus colaris LeConte, 1855: 297

    Hydroporus Clairville, 1806: 182 (Subfamily Hydroporinae, Tribe Hydroporini)
      Hydroporus americanus Aubé, 1838b: 575 - Larson et al. 2000: 293-294
      Hydroporus appalachius Sherman, 1913: 53 - MN, ON, expect in UP (Larson et al. 2000: 353-355)
        (syn.) Hydroporus signatus Sharp, 1882: 469
        (syn.) Hydroporus occidentalis Gordon and Post, 1965, not Sharp, 1882: 456
      Hydroporus badiellus Fall, 1923: 89 - UMMZ, NMNH (web page database); holotype from Marquette (Larson et al. 2000: 301-303)
      Hydroporus brevicornis Fall, 1917: 180 - UMMZ, NMNH (web page database); Larson et al. 2000: 299-300
      Hydroporus columbianus Fall, 1923: 78 - UMMZ, NMNH (web page database); Larson 1975: 294; Larson et al. 2000: 328-330
      Hydroporus dentellus Fall, 1917: 181 - UMMZ, NMNH (web page database); Larson et al. 2000: 294-296
      Hydroporus despectus Sharp, 1882: 466 - UMMZ, NMNH (web page database); see also Larson et al. 2000: 397-399
        (syn.) Hydroporus rusticus Sharp, 1882: 467
      Hydroporus dichrous Melsheimer, 1844: 28 - UMMZ, NMNH (web page database); Larson et al. 2000: 289-291
      Hydroporus fuscipennis Schaum, 1868: 64 - record at Albertson College of Idaho Collection; see also Larson et al. 2000: 345-348
        (syn.) Hydroporus puberulus Mannerheim, 1853: 163
        (syn.) Hydroporus criniticoxis Larson, 1975: 301
      Hydroporus gossei Larson and Roughley, 2000: 405 - Larson et al. 2000: 405-407
      Hydroporus larsoni Nilsson, in Nilsson & Larson, 1984: 123 - MN, ON, WI (boreal, Hilsenhoff 1995a: 10-11; Larson et al. 2000: 350-353)
        (syn.) Hydroporus fuscipennis Larson 1975: 301, not Schaum 1868
      Hydroporus melsheimeri Fall, 1917: 182 - Larson et al. 2000: 291-293
      Hydroporus morio Aubé, 1838a: 307 - Larson et al. 2000: 340-342
        (syn.) Hydroporus melanocephalus Gyllenhal, 1808: 537
        (syn.) Hydroporus caliginosus LeConte, 1850: 215
      Hydroporus niger Say, 1823b: 102 - Larson et al. 2000: 306-308
        (syn.) Hydroporus modestus Aubé, 1838b: 576
        (syn.) Hydroporus latifrons Sharp, 1882: 478
      Hydroporus nigellus Mannerheim, 1853: 163 - widespread boreal, expect in UP (Larson et al. 2000: 389-391)
        (syn.) Hydroporus sp. near despectus, Larson, 1975: 312
      Hydroporus notabilis LeConte, 1850: 216 - Larson 1975: 294; Larson et al. 2000: 303-306
        (syn.) Hydroporus arcticus Thompson, 1856: 197
        (syn.) Hydroporus niger Anderson, 1962: 62, not Say 1823: 103
      Hydroporus obscurus Sturm, 1835: 65 - Larson et al. 2000: 296-299
        (syn.) Hydroporus hockingi Larson, 1975: 296
      Hydroporus puberulus LeConte, 1850: 215 - ON, WI (boreal, Hilsenhoff 1995a: 12-13; Larson et al. 2000: 337-339)
        (syn.) Hydroporus glabriusculus Fall, 1923: 95
        (syn.) Hydroporus levanderi Sahlberg, 1888(1889): 221
      Hydroporus rectus Fall, 1923: 79 - Larson et al. 2000: 403-404
      Hydroporus rufilabris Sharp, 1882: 479 - Larson et al. 2000: 311-312
        (syn.) Hydroporus somnus Fall, 1923: 77
      Hydroporus rufinasus Mannerheim, 1852: 305 - Hilsenhoff 1995a: 13; Larson et al. 2000: 335-337
      Hydroporus signatus Mannerheim, 1853: 161 - Larson et al. 2000: 312-315
        (syn.) Hydroporus inornatus Sharp, 1882: 478
        (syn.) Hydroporus niger Sharp, 1882, not Say 1825
      Hydroporus striola (Gyllenhal in Sahlberg, 1826: 181 as Hyphydrus) - Larson et al. 2000: 365-367
        (syn.) Hydroporus subtonsus LeConte, 1855: 297
        (syn.) Hydroporus vittula Erichson, 1837: 178
      Hydroporus tartaricus LeConte, 1850: 215 - UMMZ; see also Larson et al. 2000: 385-389
        (syn.) Hydroporus elusivus Gordon, 1981: 121
      Hydroporus tenebrosus LeConte, 1850: 215 - Larson et al. 2000: 395-397
        (syn.) Hydroporus luridipennis LeConte, 1850: 216
        (syn.) Hydroporus lutulentus LeConte, 1855: 292-293
      Hydroporus tristis (Paykull, 1798: 232 as Dytiscus) - Larson et al. 2000: 343-345
        (syn.) Hydroporus varians LeConte, 1850: 215
        (syn.) Hydroporus ruficapillus Mannerheim, 1852: 304

    Hydrovatus Motschulsky, 1853: 4 (Subfamily Hydroporinae, Tribe Hydrovatini)
      Hydrovatus pustulatus (Melsheimer, 1844: 29 as Hygrotus) - Larson et al. 2000: 94-95; see Biström 1997 if other taxa in MI, or Young 1956, 1963
        (syn.) Hydrovatus indianensis Blatchley, 1910: 212

    Hygrotus Stephens, 1828: 46 (Subfamily Hydroporinae, Tribe Hydroporini)
      Hygrotus acaroides (LeConte, 1855: 294 as Hydroporus) - NMNH (web page database); see also Larson et al. 2000: 150-152
        (comb.) Coelambus acaroides winnipeg Wallis, 1926: 90
      Hygrotus compar (Fall, 1919: 8 as Coelambus) - Larson et al. 2000: 167-169
      Hygrotus dissimilis (Gemminger and Harold, 1868: 432 as Hydroporus)- Larson et al. 2000: 171-173
        (syn.) Hydroporus dispar LeConte, 1850: 216
      Hygrotus falli (Wallis, 1924: 107 as Coelambus) - Wallis 1924: 107 (Beaver Island, Charlevoix Co.); Anderson 1983: 190 (Marquette Co.); Hilsenhoff 1994: 282; Larson et al. 2000: 180-182
      Hygrotus farctus (LeConte, 1855: 294 as Hydroporus) - Larson et al. 2000: 152-153
      Hygrotus impressopunctatus (Schaller, 1783: 312 as Dytiscus) - Anderson 1983: 190-191; Larson et al. 2000: 194-196
        (syn.) Dytiscus flavicans Müller, 1776:74
        (syn.) Dytiscus lineatus Müller, 1776:74
        (syn.) Dytiscus ovalis Thunberg, 1784: 19
        (syn.) Dytiscus picipes Fabricius, 1787: 192
        (syn.) Hyphydrus lineelus Gyllenhal, 1808: 529
        (syn.) Hydroporus alternans Kunze, 1818:62
        (syn.) Hydroporus similis Kirby 1837: 68
        (syn.) Hydroporus decemlineatus Mannerheim, 1853:161.
      Hygrotus laccophilinus (LeConte, 1878: 595 as Hydroporus) - LeConte 1878a: 595; UMMZ; record at Albertson College of Idaho Collection; Larson et al. 2000: 157-159
      Hygrotus marklini (Gyllenhal, 1813: 689 as Hyphydrus) - Anderson 1983: 183 (as H. canadensis); Hilsenhoff 1994: 283; Larson et al. 2000: 162-164
        (syn.) Coelambus canadensis Fall, 1919: 11
        (syn.) Coelambus virgo Fall, 1919: 12
      Hygrotus nubilus (LeConte, 1855: 298 as Hydroporus) - Anderson 1983: 189; Larson et al. 2000: 178-180
      Hygrotus patruelis (LeConte, 1855: 298 as Hydroporus) - UMMZ; see also Anderson 1983: 194; Larson et al. 2000: 176-178
        (syn.) Hydroporus discoideus LeConte, 1855: 299
        (syn.) Coelambus borealis Fall, 1927: 137, 138
        (syn.) Coelambus quebecensis Brown, 1930b: 234
      Hygrotus picatus (Kirby, 1837: 68 as Hydroporus) - Anderson 1983: 191-192; Larson et al. 2000: 196-198
      Hygrotus punctilineatus (Fall, 1919: 17 as Coelambus) - questionable UMMZ (check); see Larson et al. 2000: 186-188
      Hygrotus sayi Balfour-Browne, 1944: 347 - Larson et al. 2000: 154-156
        (syn.) Laccophilus punctatus Say, 1824: 271 (type lost)
        (syn.) Hydroporus inaqualis Crotch, 1873: 387, not Fabricius 1777
      Hygrotus sellatus (LeConte, 1866: 365 as Hydroporus) - Anderson 1983: 182-183; Hilsenhoff 1994: 285; Larson et al. 2000: 164-166
      Hygrotus suturalis (LeConte, 1850: 216 as Hydroporus) - type specimen from Eagle Harbor, MI; Anderson 1983: 182; Hilsenhoff 1994: 285; see also Larson et al. 2000: 161-162
        (syn.) Hydroporus ovoideus LeConte, 1850: 216
      Hygrotus sylvanus (Fall, 1917: 169 as Coelambus) - UMMZ; see also Larson et al. 2000: 156-157
      Hygrotus turbidus (LeConte, 1855: 298 as Hydroporus) - Larson et al. 2000: 159-161
      Hygrotus unguicularis (Crotch, 1874: 73 as Hydroporus) - MB, QB, widespread boreal, expect in UP (Larson et al. 2000: 192-194)

    Ilybius Erichson, 1832: 18, 34 (Subfamily Colymbetinae, Tribe Agabini)
      Ilybius angustior (Gyllenhal, 1808: 500 as Dytiscus) - Larson 1987: 382-387; Larson et al. 2000: 718-720
      Ilybius biguttulus (Germar, 1824: 29 as Dytiscus) - Larson 1987: 367-371; Larson et al. 2000: 710-712; Merritt et al. 2002: 155
        (syn.) Ilybius laramaeus LeConte, 1859a: 4
      Ilybius confusus Aubé, 1838b: 280 - Larson 1987: 376-379; Larson et al. 2000: 715-716
        (syn.) Ilybius denikei Wallis 1933: 271
      Ilybius discendens Sharp, 1882: 557 - Larson 1987: 350, 360-362; Larson et al. 2000: 704-706
      Ilybius fraterculus LeConte, 1862: 521 - Larson 1987: 364-367; Hilsenhoff 1993c: 194; Larson et al. 2000: 708-709
      Ilybius ignarus (LeConte, 1862: 521 as Colymbetes) - Larson et al. 2000: 712-713
      Ilybius incarinatus Zimmermann, 1928: 181 - Larson 1987: 363, 373-374, 376; Hilsenhoff 1993c: 195; Larson et al. 2000: 713-715
      Ilybius oblitus Sharp, 1882: 560 - Larson 1987: 379-382; Hilsenhoff 1993c: 195; Larson et al. 2000: 716-717
      Ilybius picipes (Kirby, 1837: 17 as Colymbetes) - NMNH (web page database); Larson 1987: 388-391 (Isle Royale); Larson et al. 2000: 722-723
      Ilybius pleuriticus LeConte, 1850: 213 - Larson 1987: 393-396; Larson et al. 2000: 725-726
        (syn.) Ilybius inversus Sharp, 1882: 552
      Ilybius subaeneus Erichson, 1837: 156 - UMMZ; see also Larson 1987: 396-399; Larson et al. 2000: 726-728
        (syn.) Ilybius virianeus Crotch, 1873: 411
        (syn.) Ilybius suffusus Crotch, 1873: 411

    Laccophilus Leach, 1817: 69 (Subfamily Laccophilinae)
      Laccophilus biguttatus Kirby, 1837: 69 - Larson et al. 2000: 67-68
        (syn.) Laccophilus inconspicuus Fall, 1917: 164
      Laccophilus fasciatus Aubé, 1838b: 423 - Hilsenhoff 1992: 67
        (syn.) Laccophilus rufus Melsheimer, 1844: 28
        (syn.) Laccophilus terminalis Sharp, 1882: 292
      Laccophilus maculosus Say, 1823b: 100 - Larson et al. 2000: 64-65
        (syn.) Laccophilus decipiens LeConte, 1852: 205
        (syn.) Laccophilus shermani Leech, 1944: 4
      Laccophilus proximus Say, 1823b: 101 - Larson et al. 2000: 66-67
      Laccophilus undatus Aubé, 1838b: 435 - IL, IN, OH, ON, WI (Hilsenhoff 1992: 67-68; see Zimmerman 1970 if in MI

    Laccornis Gozis, 1914: 111 (Subfamily Hydroporinae, Tribe Laccornini)
      Laccornis conoideus (LeConte, 1850: 216 as Hydroporus) - LeConte 1850: 216 (Eagle Harbor, Keweenaw Co.); Wolfe and Roughley 1990: 312-313, 343; Larson et al. 2000: 79-81
        (comb.) Agaporus conoideus Zimmermann 1919: 192
        (syn.) Hydroporus oblongus Sharp, 1882: 485, not Stephens
      Laccornis deltoides (Fall, 1923: 123 as Agaporus) - Wolfe and Roughley 1990: 316-317, 343; Larson et al. 2000: 84-85
      Laccornis latens (Fall, 1937: 10 as Agaporus) - Wolfe and Roughley 1990: 315-316, 342 (Douglas Lake, Emmet Co.); Larson et al. 2000: 82-84

    Liodessus Guignot, 1939: 53 (Subfamily Hydroporinae, Tribe Bidessini)
      Liodessus affinis (Say, 1823b: 104 as Hydroporus) - Miller 1998: 299-301; Larson et al. 2000: 113-114
        (comb.) Bidessus affinis (Say, 1823) Sharp 1882: 318 and authors
      Liodessus cantralli (Young, 1953a: 111 as Bidessus) - Larson and Roughley 1990: 240; Hilsenhoff 1994: 288; Miller 1998: 310; Larson et al. 2000: 116-117
      Liodessus flavicollis (LeConte, 1855: 291 as Hydroporus) - Larson et al. 2000: 111-113; see also Miller 1998: 311
      Liodessus fuscatus (Crotch, 1873: 391 as Hydroporus) - Larson and Roughley 1990: 238-239; Larson et al. 2000: 119-120
      Liodessus obscurellus (LeConte, 1852: 206 as Hydroporus) - IN, MN, WI (Miller 1998: 303-308; Larson et al. 2000: 115-116)
        (syn.) Hydroporus macularis LeConte, 1852: 206
        (syn.) Hydroporus erythrostomus Mannerheim, 1852: 305
        (syn.) Hydroporus charlotti Clark, 1862: 182
        (syn.) Hydroporus emilianus Clark, 1862: 183
        (syn.) Bidessus nigrinus Casey, 1884: 80
        (syn.) Bidessus affinis microreticulatus (Hatch, 1828: 219) - see Miller 1998: 303-305 for taxonomic history)

    Lioporeus Guignot, 1950: 101 (Subfamily Hydroporinae, Tribe Hydroporini)
      Lioporeus triangularis (Fall, 1917: 170) - OH, NY, WI (Hilsenhoff 1994: 290; Larson et al. 2000: 240-241)

    Matus Aubé, 1836: 42 (Subfamily Colymbetinae, Tribe Matini)
      Matus bicarinatus (Say, 1823b: 98 as Colymbetes) - Larson et al. 2000: 741
      Matus ovatus Leech, 1941: 79 - Larson et al. 2000: 739-741

    Nebrioporus Régimbart, 1906: 237 (Subfamily Hydroporinae, Tribe Hydroporini)
      Nebrioporus rotundatus (LeConte, 1863: 21 as Hydroporus) - Larson et al. 2000: 450-452

    Neoporus Guignot, 1931: 46 (Subfamily Hydroporinae, Tribe Hydroporini)
      Neoporus carolinus (Fall, 1917: 180 as Hydroporus) - IN, ON (Larson et al. 2000: 260-261)
      Neoporus clypealis (Sharp, 1882: 441 as Hydroporus) Larson et al. 2000: 258-259
      Neoporus dimidiatus (Gemminger and Harold, 1868: 432 as Hydroporus) - ON, widespread (Larson et al. 2000: 266-268)
        (syn.) Hydroporus semirufus LeConte, 1850: 296
        (syn.) Hydroporus solitarius Sharp, 1882: 445
      Neoporus mellitus (LeConte, 1855: 293, 299 as Hydroporus) - Hilsenhoff 1995a: 19; Larson et al. 2000: 246-247
      Neoporus spurius (LeConte, 1855: 291, 296 as Hydroporus) - Larson et al. 2000: 255-256
      Neoporus sulcipennis (Fall, 1917: 175 as Hydroporus (orig.) - ON (Bruce Co.) Larson et al. 2000: 253-255
      Neoporus superioris (Balfour-Browne, 1944: 349 as Hydroporus) - MN, ON, WI, expect in UP (Hilsenhoff 1995a: 20; Larson et al. 2000: 268-270)
        (syn.) Hydroporus sericeus LeConte, 1850: 214
      Neoporus tennetum (Wolfe, 1984: 420 as Hydroporus) - Larson et al. 2000: 248-249
      Neoporus undulatus (Say, 1823a: 154 as Hydroporus) - UMMZ; (see also Hilsenhoff 1995a: 20-21; Larson et al. 2000: 261-264)
        (syn.) Hydroporus fasciatus Harris, 1828b: 164
        (syn.) Hydroporus pubipennis Aubé, 1838b: 480
        (syn.) Hydroporus velutinus Aubé, 1838b: 481
        (syn.) Hydroporus punctatissimus Aubé, 1838b: 484
        (syn.) Hydroporus consimilis LeConte, 1850: 214
        (syn.) Hydroporus scrutator Sharp, 1882: 440
        (syn.) Hydroporus anticus Sharp, 1882: 441
        (syn.) Hydroporus placatus Fall, 1923: 177
      Neoporus vitiosus (LeConte, 1855: 297 as Hydroporus) - Larson et al. 2000: 251-253
      Neoporus vittatus (LeConte, 1855: 296 as Hydroporus) - IL, IN, ON (western), MN, WI (Hilsenhoff 1995a: 21; Larson et al. 2000: 264-266)

    Neoscutopterus Balfour-Browne, 1943: 172 (Subfamily Colymbetinae, Tribe Colymbetini)
      Neoscutopterus angustus (LeConte, 1850: 213 as Agabus) - UMMZ, NMNH (web page database); Larson et al. 2000: 744-745
      Neoscutopterus hornii (Crotch, 1873: 405 as Scutopterus) - UMMZ, NMNH (web page database); Larson et al. 2000: 745746

    Oreodytes Seidlitz, 1887: 57 (Subfamily Hydroporinae, Tribe Hydroporini)
      Oreodytes laevis (Kirby, 1837: 67 Hydroporus) - widespread northern, may occur in UP (Larson et al. 2000: 460-461)
        (syn.) Hydroporus duodecimlineatus LeConte, 1850: 214
        (syn.) Hydroporus alpinus Crotch, 1873: 391, not Paykull, 1798
        (syn.) Hydroporus semiclarus Fall, 1923: 113
        (syn.) Hydroporus yukonensis Fall, 1926: 138.
        (syn.) Hydroporus recticollis Hatch, 1928: 220, not Fall, 1926
        (syn.) Hydroporus hortense Hatch, 1933: 27
      Oreodytes scitulus (LeConte, 1855: 295 as Hydroporus) - NMNH (web page database); see also Hilsenhoff 1994: 290; Larson et al. 2000: 466-468
        (syn.) Hydroporus septentrionalis auctorum not Gyllenhal, 1827

    Rhantus Dejean, 1833: 54 (Subfamily Colymbetinae, Tribe Colymbetini)
      Rhantus binotatus (Harris, 1828b: 164 as Colymbetes) - Zimmerman and Smith 1975: 58-68; Larson et al. 2000: 755-756
        (syn.) Colymbetes assimilis Kirby, 1837: 72
        (syn.) Colymbetes divisus Aubé, 1838b: 248
        (syn.) Rhantus flavogriseus Crotch, 1873: 409
        (syn.) Rhantus longipes Sharp, 1882: 612
        (syn.) Rhantus plejibus Sharp, 1882: 613
        (syn.) Rhantus obscurus Sharp, 1882: 613
        (syn.) Rantus aequalis Hatch, 1953: 231
      Rhantus consimilis Motschulsky, 1859: 168 - Larson et al. 2000: 752-753
        (syn.) Colymbetes tostus LeConte, 1866: 366
        (comb.syn.) Rhantus tostus (LeConte, 1866) of authors
        (syn.) Rhantus discedens Sharp, 1882: 615
      Rhantus gutticollis (Say, 1834: 442 as Colymbetes) - WI, maybe w. UP (Larson et al. 2000: 756-757)
        (syn.) Colymbetes mexicanus Laporte (de Castelnau), 1835: 101
        (syn.) Rhantus mexicanus Sharp, 1882: 614
        (syn.) Rhantus flavogriseus, Leech, 1948: 410
        (syn.) Rhantus hubbelli Hatch, 1928: 224
        (syn.) Colymbetes maculiocollis Aubé, 1838b: 245
        (syn.) Rhantus dominguensis Gemminger & Harold, 1868: 448
        (syn.) Rhantus hoppingi Wallis, 1933: 273
      Rhantus sericans Sharp, 1882: 619 - MN, ON, WI, expect in UP (Zimmerman and Smith 1975: 74-81 (as R. frontalis); Larson et al. 2000: 754-756)
      Rhantus sinuatus (LeConte, 1862: 522 as Colymbetes) - Zimmerman and Smith 1975: 105-108; Larson et al. 2000: 751-753
      Rhantus suturellus (Harris, 1828b: 164 as Colymbetes) - Zimmerman and Smith 1975: 93-100; Larson et al. 2000: 759-760
        (syn.) Rhantus bistriatus (auct. Bergstrasser, 1778: 42), of authors
        (syn.) Rhantus zimmermanni Wallis, 1933: 274
      Rhantus wallisi Hatch, 1953: 233 - Larson et al. 2000: 758-759
        (syn.) Rhantus suturellus Wallis (nec Harris), 1933: 274
        (syn.) Rhantus bistriatus Bergstrasser, F. Balfour-Browne, 1950

    Sanfilippodytes Franciscolo, 1979: 233 (Subfamily Hydroporinae, Tribe Hydroporini)
      Sanfilippodytes pseudovilis (Young, 1953b: 116 as Hydroporus) - holotype from E.S. George Reserve, Livingston Co.; Young 1953; see also Hilsenhoff 1995a: 21-22

    Stictotarsus Zimmermann, 1919: 184 (Subfamily Hydroporinae, Tribe Hydroporini)
      Stictotarsus griseostriatus (DeGeer, 1774: 403 as Dytiscus) - UMMZ, NMNH (web page database); see also Zimmerman and Smith 1975: 657-661; Larson et al. 2000: 442-444
        (comb.) Hydroporus griseostriatus, Aubé, 1838b: 541 and authors
        (syn.) Hydroporus catascopium Say, 1823b: 103
        (syn.) Hydroporus parallelus Say, 1823a: 153
        (syn). Hydroporus interruptus Say, 1834: 445 (type lost)
        (syn.) Deronectes suffusus Sharp, 1882: 434
        (syn.) Deronectes prosternalis Sharp, 1882: 434
        (syn.) Hydroporus coloradensis Fall, 1923: 108
        (syn.) Hydroporus mathiasi Hatch, 1933: 22

    Thermonectus Dejean, 1833: 61 (Subfamily Dytiscinae)
      Thermonectus basillaris (Harris, 1829: 1 as Dytiscus) - IL, IN, ON, WI (Hilsenhoff 1993a: 52; Larson et al. 2000: 826-827)
      Thermonectus nigrofasciatus ornaticollis Aubé, 1838b: 140 - WI, may occur in s. MI (Hilsenhoff 1993a: 52 (as T. ornaticollis)

    Uvarus Guignot, 1939: 53 (Subfamily Hydroporinae, Tribe Bidessini)
      Uvarus granarius (Aubé, 1838b: 501 as Hydroporus) - NMNH (web page database); see also Larson et al. 2000: 129-131
      Uvarus falli (Young, 1940: 30 as Bidessus) - FL, ON (Larson et al. 2000: 131-132)
      Uvarus lacustrus (Say, 1823b: 103 as Hydroporus) - UMMZ, NMNH (web page database); see also Larson et al. 2000: 133-134

References - Distribution

    Anderson RD. 1983. Revision of the Nearctic species of Hygrotus groups IV, V, and VI (Coleoptera: Dytiscidae). Annals of the Entomological Society of America 76(2): 173-196.
    Brown HP. 1983. A catalog of the Coleoptera of America north of Mexica. Family: Elmidae. United States Department of Agriculture Handbook 529-50. x + 23 pp.
    Downie NM, Arnett RH. 1996. The beetles of Northeastern North America. Volume I. Introduction; Suborders Archostemata, Adephaga, and Polyphaga, thru Superfamily Cantharoidea. The Sandhill Crane Press: Gainesville, Florida. xiv + 880 pp.
    Hilsenhoff WL. 1975. Notes on Nearctic Acilius (Dytiscidae), with the description of a new species. Annals of the Entomological Society of America 68(2): 271-274.
    Hilsenhoff WL. 1980. Coptotomus (Coleoptera: Dytiscidae) in eastern North America with descriptions of two new species. Transactions of the American Entomological Society 105:461-471.
    Hilsenhoff WL. 1992. Dytiscidae and Noteridae of Wisconsin (Coleoptera). I. Introduction, key to genera of adults, and distribution, habitat, life cycle, and identification of species of Abagetinae, Laccophilinae and Noteridae. The Great Lakes Entomologist 25(2): 57-69.
    Hilsenhoff WL. 1993a. Dytiscidae and Noteridae of Wisconsin (Coleoptera). II. Distribution, habitat, life cycle, and identification of species of Dytiscinae. The Great Lakes Entomologist 26(1): 35-53.
    Hilsenhoff WL. 1993b. Dytiscidae and Noteridae of Wisconsin (Coleoptera). III. Distribution, habitat, life cycle, and identification of species of species of Colymbetinae, except Agabini. The Great Lakes Entomologist 26(2): 121-136.
    Hilsenhoff WL. 1993c. Dytiscidae and Noteridae of Wisconsin (Coleoptera). IV. Distribution, habitat, life cycle, and identification of species of Agabini (Colymbetinae). The Great Lakes Entomologist 26(3): 173-197.
    Hilsenhoff WL. 1994. Dytiscidae and Noteridae of Wisconsin (Coleoptera). V. Distribution, habitat, life cycle, and identification of species of Hydroporninae, except Hydroporus Clairville sensu lato. The Great Lakes Entomologist 26(4): 275-295.
    Hilsenhoff WL. 1995a. Dytiscidae and Noteridae of Wisconsin (Coleoptera). VI. Distribution, habitat, life cycle, and identification of species of Hydroporus Clairville sensu lato (Hydroporninae). The Great Lakes Entomologist 28(1): 1-23.
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References - Taxonomic

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    July 10, 2017
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    November 21, 2012
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Page created: November 01, 2002; Last update July 11, 2017 (EB)