Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

Dytiscidae (Predaceous Diving Beetles) of Michigan

The Nearctic region has been relatively well surveyed, and most of this information has been recently included in a new publication by Larson et al. (2000). This list and nomenclature is largely based on that work as well as provisional records in the UMMZ and the Smithsonian (NMNH web page database). Genera are donoted in bold green; species denoted in bold blue have been recorded in Michigan; other species denoted in bold are likely to occur based on their known distribution. You can use the Find function in your web browser to locate families, genera, and species. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated.

Photo of Bidessonotus sp. adult from Honduras, Copyright © 2006 Ethan Bright

Acilius Leach, 1817: 69, 72 (Subfamily Dytiscinae; Tribe Aciliini)
Acilius athabascae Larson, 1975: 410-412 - MN, boreal, may occur in Isle Royale NP, nw UP (Larson et al. 2000: 813-814)
Acilius fraternus (Harris, 1828a: 156 as Dytiscus) - Hilsenhoff 1975: 273; see also Larson et al. 2000: 812
Acilius mediatus (Say, 1823b: 93 as Dytiscus) - Hilsenhoff 1975: 273; see also Larson et al. 2000: 807-808
Acilius semisulcatus Aubé, 1838b: 132 - Hilsenhoff 1975: 273-274; Larson et al. 2000: 808-810
Acilius sylvanus Hilsenhoff, 1975: 271 - Hilsenhoff 1975: 271-273; Larson et al. 2000: 811-812

Agabetes Crotch, 1873: 401 (Subfamily Agabetinae)
Agabetes acuductus (Harris, 1828b: 164 as Colyembetes) - IN, ON, WI, expect in LP (Hilsenhoff 1992: 64-65; Larson et al. 2000: 56-58)
(syn.) Agabus regulosis Aubé, 1838b: 362-363

Agabus Leach, 1817: 68 (Subfamily Colymbetinae, Tribe Agabini)
Agabus aeruginosus Aubé, 1838a: 298 - Larson et al. 2000: 596-597
Agabus ambiguus (Say, 1823b: 96 as Colymbetes.) - Larson 1989: 879-883; Larson et al. 2000: 625-626
(syn.) Agabus reticulatus Aubé, 1838b: 355
(syn.) Agabus fimbriatus LeConte, 1850: 214
Agabus antennatus Leech, 1939: 217 - MN, ON, WI, widespread north (Hilsenhoff 1993c: 181; Larson et al. 2000: 580-582)

(syn.) Agabus clavatus LeConte, 1859a: 4
Agabus anthracinus Mannerheim, 1852: 304 - Larson 1989: 896-898; Larson et al. 2000: 635-637

(syn.) Agabus scapularis Mannerheim, 1852: 303
Agabus bicolor (Kirby, 1837: 70 as Colymbetes) - Larson 1991: 1269-1271, 1314; Larson et al. 2000: 679-681

(syn.) Agabus mutus Sharp, 1882: 513
Agabus bifarius (Kirby, 1837: 71 as Colymbetes) - Larson et al. 2000: 589-591
Agabus canadensis Fall, 1922: 27 - MN, ON, WI, expect in UP (Larson 1991: 1264-1266, 1313; Hilsenhoff 1993c: 183; Larson et al. 2000: 675-676)
Agabus confinis (Gyllenhal, 1808: 511 as Dytiscus) - Larson 1991: 1287-1289, 1311; Larson et al. 2000: 692-694

(nom.nud.) Agabus ovoideus LeConte, 1863: 17
(syn.) Gaurodytes ovoideus Crotch, 1873: 418
(syn.) Gaurodytes longulus LeConte, 1878: 596
Agabus confusus (Blatchley, 1910: 229 as Rhantus) - WI (MI? found in south central WI: Hilsenhoff 1993c: 183)
(syn.) Agabus ampulus Fall, 1922: 10, 12
Agabus discolor (Harris, 1828b: 164 as Colymbetes) - Larson 1991: 1272-1275, 1315; Larson et al. 2000: 683-684
Agabus disintegratus (Crotch, 1873: 416 as Gaurodytes) - UMMZ (see also Hilsenhoff 1993c: 184; Larson et al. 2000: 600-602)
Agabus erichsoni Gemminger & Harold, 1868: 454 - Larson et al. 2000: 506-508
(syn.) Agabus nigroaeneus Erichson 1837: 157, not Marsham 1802
Agabus falli (Zimmermann, 1934: 186 as Gaurodytes) - IN, MN, ON, WI (Hilsenhoff 1993c: 185; Larson et al. 2000: 616-617)
(syn.) Agabus semipunctatus Sharp, 1882: 507, not Kirby, 1837
(syn.) Agabus sharpi Fall, 1922: 19
Agabus flavovittatus Larson and Wolfe, 1998: 44 - IL, IN, OH, ON (expect in s. MI; see Larson et al. 2000: 542)
Agabus fuscipennis (Paykull, 1798: 209 as Dytiscus) - ON (boreal, may occur in UP) (Hilsenhoff 1993c: 185; Larson 1994: 155-156, 180; Larson et al. 2000: 6583-654)
(syn.) Agabus ontarionis Fall, 1922: 30
Agabus gagates Aubé, 1838b: 306 - record at NMNH (web page database); see also Larson et al. 2000: 508-509
Agabus immaturus Larson, 1991: 1263 - Larson 1991: 1263-1264, 1313; Larson et al. 2000: 673-675
Agabus inscriptus (Crotch, 1873: 422 as Gaurodytes) - Larson 1991: 1281-1284, 1316; Larson et al. 2000: 688-689
(syn.) Agabus smithi Brown, 1930a: 88
Agabus larsoni Fery and Nilsson, 1993: 98 - ON (Larson et al. 2000: 509-510)
Agabus leptapsis (LeConte, 1878: 596 as Gaurodytes) - NMNH (web page database); see also Larson 1989: 892-894; Larson et al. 2000: 632-634
Agabus obtusatus (Say, 1823b: 99-100 as Colymbetes) - IN, OH, ON (Hilsenhoff 1993c: 187; Larson et al. 2000: 585-586)
Agabus opacus Aubé, 1837: 173 - UMMZ, NMNH (web page database - as A. pseudoconfertus Wallis); Larson 1996: 631-633, 664; Larson et al. 2000: 521-523
(syn.) Agabus sharpi Jacobson 1908 (replacement for A. sibericus Sharp 1882, not J. Shalberg 1880)
(syn.) Gaurodytes mimmi Shalberg, 1875: 182
(syn.) Agabus sibericus Sharp, 1882: 519
(syn.) Agabus gelidus Fall, 1926: 142
(syn.) Agabus pseudoconfertus Wallis, 1926: 90
(syn.) Agabus sachalinensis Kamiya, 1938: 37
Agabus phaeopterus (Kirby, 1837: 70 as Colymbetes) - Larson 1991: 1260-1262, 1312; Larson et al. 2000: 671-673
Agabus punctatus Melsheimer, 1844: 27-28 - Larson et al. 2000: 594-595
Agabus punctulatus Aubé, 1838b: 332 - IL, IN, MN, ON, WI (Hilsenhoff 1993c: 188; Larson et al. 2000: 609-610)
(syn.) Gaurodytes aeneolus Crotch, 1873: 421
Agabus semipunctatus (Kirby, 1837: 69 as Colymbetes) - Larson et al. 2000: 605-606
(syn.) Agabus stridulator Sharp, 1882: 509
(syn.) Agabus stridulatus Zimmermann, 1920: 173
Agabus semivittatus LeConte, 1852: 204 - UMMZ
Agabus seriatus (Say, 1823b: 97 as Colymbetes) - LeConte 1850: 213 (as A. parallelus); Larson et al. 2000: 559-561
(syn.) Agabus striatus Aubé, 1838a: 305
(syn.) Agabus arctus Melsheimer, 1844: 27
(syn.) Agabus parallelus LeConte, 1850: 213
(syn.) Gaurodytes intersectus Crotch, 1873: 419
Agabus subfuscatus Sharp, 1882: 514 - Larson 1991: 1271-1272, 1317; Larson et al. 2000: 681-683
Agabus thomsoni (Sahlberg, 1871: 407 as Gaurodytes) - Larson 1991: 1255-1257, 1309; Larson et al. 2000: 666-668
(syn.) Agabus congener auctorum, not Thunberg 1794: 75
Agabus wasastjernae (Sahlberg, 1824: 167 as Dytiscus) - Larson 1996: 629-631, 664; Larson et al. 2000: 519-521
(syn.) Agabus kenaiensis Fall, 1926: 141
(syn.) Agabus palustris Wallis, 1926: 92

Bidessonotus Régimbart, 1895: 331 (Subfamily Hydroporinae, Tribe Bidessini)
Bidessonotus inconspicuus (LeConte, 1855: 290 as Hydroporus) - UMMZ; see also Larson et al. 2000: 105-106
Bidessonotus pulicarius (Aubé, 1838b: 494 as Hydroporus) - IN, ON (Larson et al. 2000: 107)

Celina Aubé, 1837: 218 (Subfamily Hydroporinae, Tribe Methlini)
Celina hubbelli Young, 1979: 826 - Young 1979: 826 (Lenawee Co.); Larson et al. 2000: 90-91

Colymbetes Clairville, 1806: 188 (Subfamily Colymbetinae, Tribe Colymbetini)
Colymbetes exaratus LeConte, 1862: 522 - IL, MN, WI, may occur in the UP (Zimmerman 1981: 13-16 (as C. e. exaratus), Larson et al. 2000: 770)
(syn.) Colymbetes sculptilis exaratus, Young & Severin, 1956: 79
(syn.) Colymbetes sculptilis (in part), Larson, 1975: 389, nec LeConte
(syn.) Colymbetes rugipennis Sharp, 1882: 682
Colymbetes paykulli Erichson, 1837: 149 - Zimmerman 1981: 40-41; Larson et al. 2000: 765-766
(syn.) Colymbetes longulus LeConte, 1862: 522
Colymbetes sculptilis Harris, 1829: 8 - Zimmerman 1981: 11, 18-20, 22-27; Larson et al. 2000: 767-769

Copelatus Erichson, 1832: 18, 38 (Subfamily Copelatinae)
Copelatus chevrolati Aubé, 1838b: 389 - Larson et al. 2000: 48, 50-51
(syn.) Copelatus renovatus Guignot, 1952: 170
Copelatus glyphicus (Say, 1823b: 99 as Colymbetes) - Larson et al. 2000: 51-52

Coptotomus Say, 1834: 443 (Subfamily Colymbetinae, Tribe Coptotomini)
Coptotomus longulus LeConte, 1852: 205 - Hilsenhoff 1980: 465-466 (as C. lenticus); Larson et al. 2000: 734-736
(syn.) Coptotomus lenticus Hilsenhoff, 1980: 465
Coptotomus loticus Hilsenhoff, 1980: 467 - IL, IN, ON, WI (Hilsenhoff 1980: 467-468; Larson et al. 2000: 736-737)

Cybister Curtis, 1827: 151 (Subfamily Dytiscinae, Tribe Cybistrini)
Cybister fimbriolatus (Say, 1823b: 91 as Dytiscus) - UMMZ, NMNH (web page database); see also Larson et al. 2000: 832-833

Desmopachria Babington, 1841: 16 (Subfamily Hydroporinae, Tribe Hyphydrini)
Desmopachria (Desmopachria) convexa (Aubé, 1838b: 479 as Hydroporus) - UMMZ, NMNH (web page database); Larson et al. 2000: 99-100; see also Young 1981: 3; Hilsenhoff 1994: 277

Dytiscus Linnaeus, 1758: 411 (Subfamily Dytiscinae, Tribe Dytiscini)
Dytiscus alaskanus Balfour-Browne, 1944: 77 - MN, ON (near Sault Ste. Marie), WI, expect in UP and n. LP (Roughley 1990: 481-483, 500; Hilsenhoff 1993a: 41; Larson et al. 2000: 792-793)
(syn.) Dytiscus parvulus Motschulsky, 1852: 77
(misid.) Dytiscus ooligbuckii J. Balfour-Browne 1948: 161, Hatch 1953: 238, Kirby 1837: 74
Dytiscus carolinus Aubé, 1838b: 120 - Roughley 1990: 446-447, 453
(syn.) Dytiscus fasciventris auctorim, nec Say, 1824: 270
(syn.) ?Dytiscus fasciventris, Folkerts, 1978: 346
(syn.) Dytiscus sp., Young, 1954: 26
Dytiscus circumcinctus Ahrens, 1811: 55 - Roughley 1990: 477-481, 498; Larson et al. 2000: 791-792
(syn.) Dytiscus flavocinctus Hummel, 1823: 17
(syn.) Dytiscus angustatus Stephens, 1828: 88
(syn.) Dytiscus dubius Gyllenhal, 1827: 372
(syn.) Dytiscus ooligbuckii Kirby, 1837: 74
(syn.) Dytiscus circumscriptus Boisduval & Lacordaire, 1835: 300
(syn.) Dytiscus anxius Mannerheim, 1843: 218
(syn.) Dytiscus fusco-striatus Motschulsky, 1853: 167
(syn.) Dytiscus confusus Motschulsky, 1860: 101
(misid.) Dytiscus albinicus auctorum, nec Motschulsky, 1859: 166
(misid.) Dytiscus dauricus Anderson 1962: 73, nec Gebler, 1832: 39
Dytiscus cordieri Aubé, 1838b: 108 - Roughley 1990: 456-458, 463; Larson et al. 2000: 789-790
(syn.) Dytiscus sublimbatus LeConte, 1857: 34
(misid.) Dytiscus marginalis auctorum, nec Linnaeus 1758: 411
Dytiscus dauricus Gebler, 1832: 39 - Roughley 1990: 483-486, 501; Larson et al. 2000: 793-795
(syn.) Dytiscus confluens Say, 1834: 440 (type lost)
(syn.) Dytiscus franklinii Kirby, 1837: 77
(nom.nud.) Dytiscus frontalis Motschulsky, 1859: 489
(syn.) Dytiscus vexatus Sharp, 1882: 643
(syn.) Dytiscus dauricus var. obscurus Gschwendtner, 1922: 93
(syn.) Dytiscus amurensis J. Balfour-Browne 1944: 356 (new name for D. frontalis Motschulsky 1860 nee Marsham 1802)
Dytiscus fasciventris Say, 1824: 270 - Roughley 1990: 447-449, 454; Larson et al. 2000: 786-788
Dytiscus harrisii Kirby, 1837: 76 - Roughley 1990: 433-434, 440; Larson et al. 2000: 782-783
Dytiscus hybridus Aubé, 1838b: 116 - Roughley 1990: 434-435, 441; Larson et al. 2000: 783-785
(syn.) Leionotus compar Melsheimer, 1844: 26
Dytiscus verticalis Say, 1823b: 92 - Roughley 1990: 431-433, 439; Larson et al. 2000: 780-782

Graphoderus Dejean, 1833: 61 (Subfamily Dytiscinae, Tribe Aciliini)
Graphoderus fasciocollis (Harris, 1828a: 156 as Dytiscus) - Larson et al. 2000: 821-822
Graphoderus liberus (Say, 1825: 160 as Dytiscus) - Larson et al. 2000: 817-818
(syn.) Dytiscus thoracicus Harris, 1828a: 156
(syn.) Colymbetes rugicollis Kirby, 1837: 73
(syn.) Hydaticus brunnipennis Aubé, 1838b: 203
Graphoderus manitobensis Wallis, 1933: 276 - IL, WI, expect in western UP (Hilsenhoff 1993a: 47-48; Larson et al. 2000: 822-823)
Graphoderus occidentalis Horn, 1883: 281 - NMNH (web page database); see also Larson et al. 2000: 820-821
Graphoderus perplexus Sharp, 1882: 695 - Larson et al. 2000: 818-819
(syn.) Graphoderus elatus Sharp, 1882: 695
(misid.) Graphoderus cinereus Horn, 1883: 280 (not Linnaeus, 1758)
(misid.) Graphoderus zonatus Zimmermann 1919: 231 (not Hoppe, 1795)
(misid.) Graphoderus zonatus Hatch 1928 (not Hoppe, 1795)

Heterosternuta Strand, 1935: 156, 161, 178 (Subfamily Hydroporinae, Tribe Hydroporini)
Heterosternuta pulcher (LeConte, 1855: 298 as Hydroporus) - OH, ON, MN (Hilsenhoff 1995a: 4; Larson et al. 2000: 235-236)
Heterosternuta wickhami (Zaitzev, 1908: 118 as Hydroporus) - UMMZ; see also Larson et al. 2000: 232-234

Hydaticus Leach, 1817: 69 (Subfamily Dytiscinae, Tribe Hydaticini)
Hydaticus aruspex Clark, 1864: 212 - Roughley and Pengelly 1981: 260-267, 300 (as H. modestus); Larson et al. 2000: 800-801
(syn.) Hydaticus modestus Sharp, 1882: 650
Hydaticus piceus LeConte, 1863: 23 - Roughley and Pengelly 1981: 267-268, 302; Larson et al. 2000: 802-803

Hydrocolus (=Hydroporus oblitus-group Fall, 1923) Roughley & Larson, in Larson, Alarie & Roughley 2000: 407 (Subfamily Hydroporinae, Tribe Hydroporini)
Hydrocolus paugus (Fall, 1923: 50 as Hydroporus) - Larson et al. 2000: 414-416
Hydrocolus persimilis (Crotch, 1873: 395 as Hydroporus) - Larson et al. 2000: 416-417
(syn.) Hydroporus aequus Fall, 1923: 53
(syn.) Hydroporus acadianus J Balfour-Brown, 1948: 163
Hydrocolus rybyae (Larson, 1975: 290 as Hydroporus) - ON, WI (boreal, expect in UP; Hilsenhoff 1995a: 16; Larson et al. 2000: 422-423)
Hydrocolus stagnalis (Gemminger & Harold, 1868: 441 as Hydroporus (orig.) - Larson et al. 2000: 426-428
(syn.) Hydroporus colaris LeConte, 1855: 297

Hydroporus Clairville, 1806: 182 (Subfamily Hydroporinae, Tribe Hydroporini)
Hydroporus americanus Aubé, 1838b: 575 - Larson et al. 2000: 293-294
Hydroporus appalachius Sherman, 1913: 53 - MN, ON, expect in UP (Larson et al. 2000: 353-355)
(syn.) Hydroporus signatus Sharp, 1882: 469
(syn.) Hydroporus occidentalis Gordon and Post, 1965, not Sharp, 1882: 456

Hydroporus badiellus Fall, 1923: 89 - UMMZ, NMNH (web page database); holotype from Marquette (Larson et al. 2000: 301-303)
Hydroporus brevicornis Fall, 1917: 180 - UMMZ, NMNH (web page database); Larson et al. 2000: 299-300
Hydroporus columbianus Fall, 1923: 78 - UMMZ, NMNH (web page database); Larson 1975: 294; Larson et al. 2000: 328-330
Hydroporus dentellus Fall, 1917: 181 - UMMZ, NMNH (web page database); Larson et al. 2000: 294-296
Hydroporus despectus Sharp, 1882: 466 - UMMZ, NMNH (web page database); see also Larson et al. 2000: 397-399
(syn.) Hydroporus rusticus Sharp, 1882: 467
Hydroporus dichrous Melsheimer, 1844: 28 - UMMZ, NMNH (web page database); Larson et al. 2000: 289-291
Hydroporus fuscipennis Schaum, 1868: 64 - record at Albertson College of Idaho Collection; see also Larson et al. 2000: 345-348

(syn.) Hydroporus puberulus Mannerheim, 1853: 163
(syn.) Hydroporus criniticoxis Larson, 1975: 301
Hydroporus gossei Larson and Roughley, 2000: 405 - Larson et al. 2000: 405-407
Hydroporus larsoni Nilsson, in Nilsson & Larson, 1984: 123 - MN, ON, WI (boreal, Hilsenhoff 1995a: 10-11; Larson et al. 2000: 350-353)

(syn.) Hydroporus fuscipennis Larson 1975: 301, not Schaum 1868
Hydroporus melsheimeri Fall, 1917: 182 - Larson et al. 2000: 291-293
Hydroporus morio Aubé, 1838a: 307 - Larson et al. 2000: 340-342

(syn.) Hydroporus melanocephalus Gyllenhal, 1808: 537
(syn.) Hydroporus caliginosus LeConte, 1850: 215
Hydroporus niger Say, 1823b: 102 - Larson et al. 2000: 306-308
(syn.) Hydroporus modestus Aubé, 1838b: 576
(syn.) Hydroporus latifrons Sharp, 1882: 478
Hydroporus nigellus Mannerheim, 1853: 163 - widespread boreal, expect in UP (Larson et al. 2000: 389-391)
(syn.) Hydroporus sp. near despectus, Larson, 1975: 312
Hydroporus notabilis LeConte, 1850: 216 - Larson 1975: 294; Larson et al. 2000: 303-306
(syn.) Hydroporus arcticus Thompson, 1856: 197
(syn.) Hydroporus niger Anderson, 1962: 62, not Say 1823: 103
Hydroporus obscurus Sturm, 1835: 65 - Larson et al. 2000: 296-299
(syn.) Hydroporus hockingi Larson, 1975: 296
Hydroporus puberulus LeConte, 1850: 215 - ON, WI (boreal, Hilsenhoff 1995a: 12-13; Larson et al. 2000: 337-339)
(syn.) Hydroporus glabriusculus Fall, 1923: 95
(syn.) Hydroporus levanderi Sahlberg, 1888(1889): 221
Hydroporus rectus Fall, 1923: 79 - Larson et al. 2000: 403-404
Hydroporus rufilabris Sharp, 1882: 479 - Larson et al. 2000: 311-312
(syn.) Hydroporus somnus Fall, 1923: 77
Hydroporus rufinasus Mannerheim, 1852: 305 - Hilsenhoff 1995a: 13; Larson et al. 2000: 335-337
Hydroporus signatus Mannerheim, 1853: 161 - Larson et al. 2000: 312-315
(syn.) Hydroporus inornatus Sharp, 1882: 478
(syn.) Hydroporus niger Sharp, 1882, not Say 1825
Hydroporus striola (Gyllenhal in Sahlberg, 1826: 181 as Hyphydrus) - Larson et al. 2000: 365-367
(syn.) Hydroporus subtonsus LeConte, 1855: 297
(syn.) Hydroporus vittula Erichson, 1837: 178
Hydroporus tartaricus LeConte, 1850: 215 - UMMZ; see also Larson et al. 2000: 385-389
(syn.) Hydroporus elusivus Gordon, 1981: 121
Hydroporus tenebrosus LeConte, 1850: 215 - Larson et al. 2000: 395-397
(syn.) Hydroporus luridipennis LeConte, 1850: 216
(syn.) Hydroporus lutulentus LeConte, 1855: 292-293
Hydroporus tristis (Paykull, 1798: 232 as Dytiscus) - Larson et al. 2000: 343-345
(syn.) Hydroporus varians LeConte, 1850: 215
(syn.) Hydroporus ruficapillus Mannerheim, 1852: 304

Hydrovatus Motschulsky, 1853: 4 (Subfamily Hydroporinae, Tribe Hydrovatini)
Hydrovatus pustulatus (Melsheimer, 1844: 29 as Hygrotus) - Larson et al. 2000: 94-95; see Biström 1997 if other taxa in MI, or Young 1956, 1963
(syn.) Hydrovatus indianensis Blatchley, 1910: 212

Hygrotus Stephens, 1828: 46 (Subfamily Hydroporinae, Tribe Hydroporini)
Hygrotus acaroides (LeConte, 1855: 294 as Hydroporus) - NMNH (web page database); see also Larson et al. 2000: 150-152
(comb.) Coelambus acaroides winnipeg Wallis, 1926: 90
Hygrotus compar (Fall, 1919: 8 as Coelambus) - Larson et al. 2000: 167-169
Hygrotus dissimilis (Gemminger and Harold, 1868: 432 as Hydroporus)- Larson et al. 2000: 171-173
(syn.) Hydroporus dispar LeConte, 1850: 216
Hygrotus falli (Wallis, 1924: 107 as Coelambus) - Wallis 1924: 107 (Beaver Island, Charlevoix Co.); Anderson 1983: 190 (Marquette Co.); Hilsenhoff 1994: 282; Larson et al. 2000: 180-182
Hygrotus farctus (LeConte, 1855: 294 as Hydroporus) - Larson et al. 2000: 152-153
Hygrotus impressopunctatus (Schaller, 1783: 312 as Dytiscus) - Anderson 1983: 190-191; Larson et al. 2000: 194-196
(syn.) Dytiscus flavicans Müller, 1776:74
(syn.) Dytiscus lineatus Müller, 1776:74
(syn.) Dytiscus ovalis Thunberg, 1784: 19
(syn.) Dytiscus picipes Fabricius, 1787: 192
(syn.) Hyphydrus lineelus Gyllenhal, 1808: 529
(syn.) Hydroporus alternans Kunze, 1818:62
(syn.) Hydroporus similis Kirby 1837: 68
(syn.) Hydroporus decemlineatus Mannerheim, 1853:161.
Hygrotus laccophilinus (LeConte, 1878: 595 as Hydroporus) - LeConte 1878a: 595; UMMZ; record at Albertson College of Idaho Collection; Larson et al. 2000: 157-159
Hygrotus marklini (Gyllenhal, 1813: 689 as Hyphydrus) - Anderson 1983: 183 (as H. canadensis); Hilsenhoff 1994: 283; Larson et al. 2000: 162-164
(syn.) Coelambus canadensis Fall, 1919: 11
(syn.) Coelambus virgo Fall, 1919: 12
Hygrotus nubilus (LeConte, 1855: 298 as Hydroporus) - Anderson 1983: 189; Larson et al. 2000: 178-180
Hygrotus patruelis (LeConte, 1855: 298 as Hydroporus) - UMMZ; see also Anderson 1983: 194; Larson et al. 2000: 176-178
(syn.) Hydroporus discoideus LeConte, 1855: 299
(syn.) Coelambus borealis Fall, 1927: 137, 138
(syn.) Coelambus quebecensis Brown, 1930b: 234
Hygrotus picatus (Kirby, 1837: 68 as Hydroporus) - Anderson 1983: 191-192; Larson et al. 2000: 196-198
Hygrotus punctilineatus (Fall, 1919: 17 as Coelambus) - questionable UMMZ (check); see Larson et al. 2000: 186-188
Hygrotus sayi Balfour-Browne, 1944: 347 - Larson et al. 2000: 154-156
(syn.) Laccophilus punctatus Say, 1824: 271 (type lost)
(syn.) Hydroporus inaqualis Crotch, 1873: 387, not Fabricius 1777
Hygrotus sellatus (LeConte, 1866: 365 as Hydroporus) - Anderson 1983: 182-183; Hilsenhoff 1994: 285; Larson et al. 2000: 164-166
Hygrotus suturalis (LeConte, 1850: 216 as Hydroporus) - type specimen from Eagle Harbor, MI; Anderson 1983: 182; Hilsenhoff 1994: 285; see also Larson et al. 2000: 161-162
(syn.) Hydroporus ovoideus LeConte, 1850: 216
Hygrotus sylvanus (Fall, 1917: 169 as Coelambus) - UMMZ; see also Larson et al. 2000: 156-157
Hygrotus turbidus (LeConte, 1855: 298 as Hydroporus) - Larson et al. 2000: 159-161
Hygrotus unguicularis (Crotch, 1874: 73 as Hydroporus) - MB, QB, widespread boreal, expect in UP (Larson et al. 2000: 192-194)

Ilybius Erichson, 1832: 18, 34 (Subfamily Colymbetinae, Tribe Agabini)
Ilybius angustior (Gyllenhal, 1808: 500 as Dytiscus) - Larson 1987: 382-387; Larson et al. 2000: 718-720
Ilybius biguttulus (Germar, 1824: 29 as Dytiscus) - Larson 1987: 367-371; Larson et al. 2000: 710-712; Merritt et al. 2002: 155
(syn.) Ilybius laramaeus LeConte, 1859a: 4
Ilybius confusus Aubé, 1838b: 280 - Larson 1987: 376-379; Larson et al. 2000: 715-716
(syn.) Ilybius denikei Wallis 1933: 271
Ilybius discendens Sharp, 1882: 557 - Larson 1987: 350, 360-362; Larson et al. 2000: 704-706
Ilybius fraterculus LeConte, 1862: 521 - Larson 1987: 364-367; Hilsenhoff 1993c: 194; Larson et al. 2000: 708-709
Ilybius ignarus (LeConte, 1862: 521 as Colymbetes) - Larson et al. 2000: 712-713
Ilybius incarinatus Zimmermann, 1928: 181 - Larson 1987: 363, 373-374, 376; Hilsenhoff 1993c: 195; Larson et al. 2000: 713-715
Ilybius oblitus Sharp, 1882: 560 - Larson 1987: 379-382; Hilsenhoff 1993c: 195; Larson et al. 2000: 716-717
Ilybius picipes (Kirby, 1837: 17 as Colymbetes) - NMNH (web page database); Larson 1987: 388-391 (Isle Royale); Larson et al. 2000: 722-723
Ilybius pleuriticus LeConte, 1850: 213 - Larson 1987: 393-396; Larson et al. 2000: 725-726
(syn.) Ilybius inversus Sharp, 1882: 552
Ilybius subaeneus Erichson, 1837: 156 - UMMZ; see also Larson 1987: 396-399; Larson et al. 2000: 726-728
(syn.) Ilybius virianeus Crotch, 1873: 411
(syn.) Ilybius suffusus Crotch, 1873: 411

Laccophilus Leach, 1817: 69 (Subfamily Laccophilinae)
Laccophilus biguttatus Kirby, 1837: 69 - Larson et al. 2000: 67-68
(syn.) Laccophilus inconspicuus Fall, 1917: 164

Laccophilus fasciatus Aubé, 1838b: 423 - Hilsenhoff 1992: 67
(syn.) Laccophilus rufus Melsheimer, 1844: 28
(syn.) Laccophilus terminalis Sharp, 1882: 292
Laccophilus maculosus Say, 1823b: 100 - Larson et al. 2000: 64-65
(syn.) Laccophilus decipiens LeConte, 1852: 205
(syn.) Laccophilus shermani Leech, 1944: 4
Laccophilus proximus Say, 1823b: 101 - Larson et al. 2000: 66-67
Laccophilus undatus Aubé, 1838b: 435 - IL, IN, OH, ON, WI (Hilsenhoff 1992: 67-68; see Zimmerman 1970 if in MI


Laccornis Gozis, 1914: 111 (Subfamily Hydroporinae, Tribe Laccornini)
Laccornis conoideus (LeConte, 1850: 216 as Hydroporus) - LeConte 1850: 216 (Eagle Harbor, Keweenaw Co.); Wolfe and Roughley 1990: 312-313, 343; Larson et al. 2000: 79-81
(comb.) Agaporus conoideus Zimmermann 1919: 192
(syn.) Hydroporus oblongus Sharp, 1882: 485, not Stephens
Laccornis deltoides (Fall, 1923: 123 as Agaporus) - Wolfe and Roughley 1990: 316-317, 343; Larson et al. 2000: 84-85
Laccornis latens (Fall, 1937: 10 as Agaporus) - Wolfe and Roughley 1990: 315-316, 342 (Douglas Lake, Emmet Co.); Larson et al. 2000: 82-84

Liodessus Guignot, 1939: 53 (Subfamily Hydroporinae, Tribe Bidessini)
Liodessus affinis (Say, 1823b: 104 as Hydroporus) - Miller 1998: 299-301; Larson et al. 2000: 113-114
(comb.) Bidessus affinis (Say, 1823) Sharp 1882: 318 and authors
Liodessus cantralli (Young, 1953a: 111 as Bidessus) - Larson and Roughley 1990: 240; Hilsenhoff 1994: 288; Miller 1998: 310; Larson et al. 2000: 116-117
Liodessus flavicollis (LeConte, 1855: 291 as Hydroporus) - Larson et al. 2000: 111-113; see also Miller 1998: 311
Liodessus fuscatus (Crotch, 1873: 391 as Hydroporus) - Larson and Roughley 1990: 238-239; Larson et al. 2000: 119-120
Liodessus obscurellus (LeConte, 1852: 206 as Hydroporus) - IN, MN, WI (Miller 1998: 303-308; Larson et al. 2000: 115-116)
(syn.) Hydroporus macularis LeConte, 1852: 206
(syn.) Hydroporus erythrostomus Mannerheim, 1852: 305
(syn.) Hydroporus charlotti Clark, 1862: 182
(syn.) Hydroporus emilianus Clark, 1862: 183
(syn.) Bidessus nigrinus Casey, 1884: 80
(syn.) Bidessus affinis microreticulatus (Hatch, 1828: 219) - see Miller 1998: 303-305 for taxonomic history)

Lioporeus Guignot, 1950: 101 (Subfamily Hydroporinae, Tribe Hydroporini)
Lioporeus triangularis (Fall, 1917: 170) - OH, NY, WI (Hilsenhoff 1994: 290; Larson et al. 2000: 240-241)

Matus Aubé, 1836: 42 (Subfamily Colymbetinae, Tribe Matini)
Matus bicarinatus (Say, 1823b: 98 as Colymbetes) - Larson et al. 2000: 741
Matus ovatus Leech, 1941: 79 - Larson et al. 2000: 739-741

Nebrioporus Régimbart, 1906: 237 (Subfamily Hydroporinae, Tribe Hydroporini)
Nebrioporus rotundatus (LeConte, 1863: 21 as Hydroporus) - Larson et al. 2000: 450-452

Neoporus Guignot, 1931: 46 (Subfamily Hydroporinae, Tribe Hydroporini)
Neoporus carolinus (Fall, 1917: 180 as Hydroporus) - IN, ON (Larson et al. 2000: 260-261)
Neoporus clypealis (Sharp, 1882: 441 as Hydroporus) Larson et al. 2000: 258-259
Neoporus dimidiatus (Gemminger and Harold, 1868: 432 as Hydroporus) - ON, widespread (Larson et al. 2000: 266-268)
(syn.) Hydroporus semirufus LeConte, 1850: 296
(syn.) Hydroporus solitarius Sharp, 1882: 445
Neoporus mellitus (LeConte, 1855: 293, 299 as Hydroporus) - Hilsenhoff 1995a: 19; Larson et al. 2000: 246-247
Neoporus spurius (LeConte, 1855: 291, 296 as Hydroporus) - Larson et al. 2000: 255-256
Neoporus sulcipennis (Fall, 1917: 175 as Hydroporus (orig.) - ON (Bruce Co.) Larson et al. 2000: 253-255
Neoporus superioris (Balfour-Browne, 1944: 349 as Hydroporus) - MN, ON, WI, expect in UP (Hilsenhoff 1995a: 20; Larson et al. 2000: 268-270)
(syn.) Hydroporus sericeus LeConte, 1850: 214
Neoporus tennetum (Wolfe, 1984: 420 as Hydroporus) - Larson et al. 2000: 248-249
Neoporus undulatus (Say, 1823a: 154 as Hydroporus) - UMMZ; (see also Hilsenhoff 1995a: 20-21; Larson et al. 2000: 261-264)
(syn.) Hydroporus fasciatus Harris, 1828b: 164
(syn.) Hydroporus pubipennis Aubé, 1838b: 480
(syn.) Hydroporus velutinus Aubé, 1838b: 481
(syn.) Hydroporus punctatissimus Aubé, 1838b: 484
(syn.) Hydroporus consimilis LeConte, 1850: 214
(syn.) Hydroporus scrutator Sharp, 1882: 440
(syn.) Hydroporus anticus Sharp, 1882: 441
(syn.) Hydroporus placatus Fall, 1923: 177
Neoporus vitiosus (LeConte, 1855: 297 as Hydroporus) - Larson et al. 2000: 251-253
Neoporus vittatus (LeConte, 1855: 296 as Hydroporus) - IL, IN, ON (western), MN, WI (Hilsenhoff 1995a: 21; Larson et al. 2000: 264-266)

Neoscutopterus Balfour-Browne, 1943: 172 (Subfamily Colymbetinae, Tribe Colymbetini)
Neoscutopterus angustus (LeConte, 1850: 213 as Agabus) - UMMZ, NMNH (web page database); Larson et al. 2000: 744-745
Neoscutopterus hornii (Crotch, 1873: 405 as Scutopterus) - UMMZ, NMNH (web page database); Larson et al. 2000: 745746

Oreodytes Seidlitz, 1887: 57 (Subfamily Hydroporinae, Tribe Hydroporini)
Oreodytes laevis (Kirby, 1837: 67 Hydroporus) - Isle Royale (Gerald Shepard and Kurt Schmude, pers.comm. 2017), widespread northern, may occur elsewhere in the UP (Larson et al. 2000: 460-461)
(syn.) Hydroporus duodecimlineatus LeConte, 1850: 214
(syn.) Hydroporus alpinus Crotch, 1873: 391, not Paykull, 1798
(syn.) Hydroporus semiclarus Fall, 1923: 113
(syn.) Hydroporus yukonensis Fall, 1926: 138.
(syn.) Hydroporus recticollis Hatch, 1928: 220, not Fall, 1926
(syn.) Hydroporus hortense Hatch, 1933: 27
Oreodytes scitulus (LeConte, 1855: 295 as Hydroporus) - NMNH (web page database); see also Hilsenhoff 1994: 290; Larson et al. 2000: 466-468
(syn.) Hydroporus septentrionalis auctorum not Gyllenhal, 1827

Rhantus Dejean, 1833: 54 (Subfamily Colymbetinae, Tribe Colymbetini)
Rhantus binotatus (Harris, 1828b: 164 as Colymbetes) - Zimmerman and Smith 1975: 58-68; Larson et al. 2000: 755-756
(syn.) Colymbetes assimilis Kirby, 1837: 72
(syn.) Colymbetes divisus Aubé, 1838b: 248
(syn.) Rhantus flavogriseus Crotch, 1873: 409
(syn.) Rhantus longipes Sharp, 1882: 612
(syn.) Rhantus plejibus Sharp, 1882: 613
(syn.) Rhantus obscurus Sharp, 1882: 613
(syn.) Rantus aequalis Hatch, 1953: 231
Rhantus consimilis Motschulsky, 1859: 168 - Larson et al. 2000: 752-753
(syn.) Colymbetes tostus LeConte, 1866: 366
(comb.syn.) Rhantus tostus (LeConte, 1866) of authors
(syn.) Rhantus discedens Sharp, 1882: 615
Rhantus gutticollis (Say, 1834: 442 as Colymbetes) - WI, maybe w. UP (Larson et al. 2000: 756-757)
(syn.) Colymbetes mexicanus Laporte (de Castelnau), 1835: 101
(syn.) Rhantus mexicanus Sharp, 1882: 614
(syn.) Rhantus flavogriseus, Leech, 1948: 410
(syn.) Rhantus hubbelli Hatch, 1928: 224
(syn.) Colymbetes maculiocollis Aubé, 1838b: 245
(syn.) Rhantus dominguensis Gemminger & Harold, 1868: 448
(syn.) Rhantus hoppingi Wallis, 1933: 273
Rhantus sericans Sharp, 1882: 619 - MN, ON, WI, expect in UP (Zimmerman and Smith 1975: 74-81 (as R. frontalis); Larson et al. 2000: 754-756)
Rhantus sinuatus (LeConte, 1862: 522 as Colymbetes) - Zimmerman and Smith 1975: 105-108; Larson et al. 2000: 751-753
Rhantus suturellus (Harris, 1828b: 164 as Colymbetes) - Zimmerman and Smith 1975: 93-100; Larson et al. 2000: 759-760
(syn.) Rhantus bistriatus (auct. Bergstrasser, 1778: 42), of authors
(syn.) Rhantus zimmermanni Wallis, 1933: 274
Rhantus wallisi Hatch, 1953: 233 - Larson et al. 2000: 758-759
(syn.) Rhantus suturellus Wallis (nec Harris), 1933: 274
(syn.) Rhantus bistriatus Bergstrasser, F. Balfour-Browne, 1950

Sanfilippodytes Franciscolo, 1979: 233 (Subfamily Hydroporinae, Tribe Hydroporini)
Sanfilippodytes pseudovilis (Young, 1953b: 116 as Hydroporus) - holotype from E.S. George Reserve, Livingston Co.; Young 1953; see also Hilsenhoff 1995a: 21-22

Stictotarsus Zimmermann, 1919: 184 (Subfamily Hydroporinae, Tribe Hydroporini)
Stictotarsus griseostriatus (DeGeer, 1774: 403 as Dytiscus) - UMMZ, NMNH (web page database); see also Zimmerman and Smith 1975: 657-661; Larson et al. 2000: 442-444
(comb.) Hydroporus griseostriatus, Aubé, 1838b: 541 and authors
(syn.) Hydroporus catascopium Say, 1823b: 103
(syn.) Hydroporus parallelus Say, 1823a: 153
(syn). Hydroporus interruptus Say, 1834: 445 (type lost)
(syn.) Deronectes suffusus Sharp, 1882: 434
(syn.) Deronectes prosternalis Sharp, 1882: 434
(syn.) Hydroporus coloradensis Fall, 1923: 108
(syn.) Hydroporus mathiasi Hatch, 1933: 22

Thermonectus Dejean, 1833: 61 (Subfamily Dytiscinae)
Thermonectus basillaris (Harris, 1829: 1 as Dytiscus) - IL, IN, ON, WI (Hilsenhoff 1993a: 52; Larson et al. 2000: 826-827)
Thermonectus nigrofasciatus ornaticollis Aubé, 1838b: 140 - WI, may occur in s. MI (Hilsenhoff 1993a: 52 (as T. ornaticollis)

Uvarus Guignot, 1939: 53 (Subfamily Hydroporinae, Tribe Bidessini)
Uvarus granarius (Aubé, 1838b: 501 as Hydroporus) - NMNH (web page database); see also Larson et al. 2000: 129-131
Uvarus falli (Young, 1940: 30 as Bidessus) - FL, ON (Larson et al. 2000: 131-132)
Uvarus lacustrus (Say, 1823b: 103 as Hydroporus) - UMMZ, NMNH (web page database); see also Larson et al. 2000: 133-134

References - Distribution

Anderson RD. 1983. Revision of the Nearctic species of Hygrotus groups IV, V, and VI (Coleoptera: Dytiscidae). Annals of the Entomological Society of America 76(2): 173-196.
Downie NM, Arnett RH. 1996. The beetles of Northeastern North America. Volume I. Introduction; Suborders Archostemata, Adephaga, and Polyphaga, thru Superfamily Cantharoidea. The Sandhill Crane Press: Gainesville, Florida. xiv + 880 pp.
Fabricius JC. 1798. Entomologia systematica. Supplement. Proft et Storch, Hafniae. [2] + 572 + 52 pp. (PDF)
Hilsenhoff WL. 1975. Notes on Nearctic Acilius (Dytiscidae), with the description of a new species. Annals of the Entomological Society of America 68(2): 271-274.
Hilsenhoff WL. 1980. Coptotomus (Coleoptera: Dytiscidae) in eastern North America with descriptions of two new species. Transactions of the American Entomological Society 105:461-471.
Hilsenhoff WL. 1992. Dytiscidae and Noteridae of Wisconsin (Coleoptera). I. Introduction, key to genera of adults, and distribution, habitat, life cycle, and identification of species of Abagetinae, Laccophilinae and Noteridae. The Great Lakes Entomologist 25(2): 57-69.
Hilsenhoff WL. 1993a. Dytiscidae and Noteridae of Wisconsin (Coleoptera). II. Distribution, habitat, life cycle, and identification of species of Dytiscinae. The Great Lakes Entomologist 26(1): 35-53.
Hilsenhoff WL. 1993b. Dytiscidae and Noteridae of Wisconsin (Coleoptera). III. Distribution, habitat, life cycle, and identification of species of species of Colymbetinae, except Agabini. The Great Lakes Entomologist 26(2): 121-136.
Hilsenhoff WL. 1993c. Dytiscidae and Noteridae of Wisconsin (Coleoptera). IV. Distribution, habitat, life cycle, and identification of species of Agabini (Colymbetinae). The Great Lakes Entomologist 26(3): 173-197.
Hilsenhoff WL. 1994. Dytiscidae and Noteridae of Wisconsin (Coleoptera). V. Distribution, habitat, life cycle, and identification of species of Hydroporninae, except Hydroporus Clairville sensu lato. The Great Lakes Entomologist 26(4): 275-295.
Hilsenhoff WL. 1995a. Dytiscidae and Noteridae of Wisconsin (Coleoptera). VI. Distribution, habitat, life cycle, and identification of species of Hydroporus Clairville sensu lato (Hydroporninae). The Great Lakes Entomologist 28(1): 1-23.
Larson DJ. 1987. Revision of North American species of Ilybius Erichson (Coleoptera: Dytiscidae), with systematic notes on Palaearctic species. Journal of the New York Entomological Society 95(3):341-413.
Larson DJ. 1989. Revision of North American Agabus Leach (Coleoptera: Dytiscidae): Introduction, key to species groups, and classification of the ambiguus-, tristus-, and arcticus-groups. The The Canadian Entomologist 121(10): 861-919.
Larson DJ. 1991. Revision of North American Agabus Leach (Coleoptera: Dytiscidae): elongatus-, zetterstedti-, and confinis-groups. The The Canadian Entomologist 123(6): 1239-1317.
Larson DJ. 1994. Revision of North American Agabus Leach (Coleoptera: Dytiscidae): lutosus-, obsoletus-, and fuscipennis-groups. The The Canadian Entomologist 126(1): 135-181.
Larson DJ. 1996. Revision of North American Agabus Leach (Coleoptera: Dytiscidae): the opacus-group. The The Canadian Entomologist 128(04): 613-665.
Larson DJ, Roughley RE. 1990. A review of the specie sof Liodessus Guignot of North America north of Mexico with the description of a new species (Coleoptera: Dytiscidae). Journal of the New York Entomological Society 98(2): 233-245.
Larson DJ., Alarie Y., Roughley RE. 2000. Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa, Ontario, Canada. 982 pp.
Merritt RW, Benbow ME, Hudson PL. 2002. Wetland macroinvertebrates of Prentiss Bay, Lake Huron, Michigan: diversity and function group comparison. The Great Lakes Entomologist 35(2): 149-160.
Miller KB. 1998. Revision of the Nearctic Liodessus affinis (Say 1823) species group (Coleoptera: Dytscidae, Hydroporinae, Bidessini). Entomologica scandinavica 29(3):281-314.
Roughley RE. 1990. A systematic revision of species of Dytiscus Linnaeus (Coleoptera: Dytiscidae). Part 1. Classification based on adult stage. Quaestiones Entomologicae 26:383-557.
Roughley RE, Pengelly DH. 1981. Classification, phylogeny, and zoogeography of Hydaticus Leach (Coleoptera: Dytiscidae) of North America. Questiones Entomologicae 17: 249-309.
Wolfe GW, Roughley RE. 1990. A taxonomic, phylogenetic, and zoogeographic analysis of Laccornis Gozis (Coleoptera: Dytiscidae) with the description of Laccornini, a new tribe of Hydroporinae. Quaestiones Entomologicae 26(3): 273-354.
Young FN. 1953. A new Hydroporus from Michigan, with notes on other members of the Hydroporus vilis group (Coleoptera: Dytiscidae). Bulletin of the Brooklyn Entomological Society 48: 115-122.
Young FN. 1979. A key to the Nearctic species of Celina with descriptions of new species (Coleoptera: Dytiscidae). Journal of the Kansas Entomological Society 52: 820-830.
Young FN. 1981. Predaceous water beetles of the genus Desmopachria: the convexa-grana group. Occasional Papers of the Florida State Collection of Arthropods 2: 1-11.
Young DK, Stribling JB. 1990. Systematics of the North American Cyphon collaris species complex with the description of a new species (Coleoptera: Scirtidae). Proceedings of the Entomological Society of Washington 92(2): 194-204.
Zimmerman JR. 1981. A revision of the Colymbetes of North America (Dytiscidae). The Coleopterists Bulletin 35(1):1-52.
Zimmerman JR, Smith AH. 1975. A survey of the Deronectes (Coleoptera: Dytiscidae) of Canada, the United States, and northern Mexico. Transactions of the American Entomological Society 101(4): 651-723.
Zimmerman JR, Smith RL. 1975. The genus Rhantus (Coleoptera: Dytiscidae) in North America. Part I. General account of the species. Transactions of the American Entomological Society 101(1): 33-123.

References - Taxonomic

Ahrens A. 1811. Beiträge zu einer Monographie der deutschen Rohrkäfer-Arten. Neue Schriften der naturforschenden Gesellschaft zu Halle 1: 43-67.
Anderson RD. 1962. The Dytiscidae (Coleoptera) of Utah: keys, original citation, types and Utah distribution. Great Basin Naturalist 22: 54-75.
Aube C. 1836-1838a. Tome cinquième. Hydrocanthares, in Dejean PFMA (ed.), Iconographie et histoire naturelle des Coléoptères d'Europe. Méquignon-Marvis, Paris 416 pp. + 1-46 pls. [pp. xi + 1-64 (1836); 65-224 (1837); 225-416 (1838)].
Aube C. 1838. Tome sixème. Species général des coléoptères, de la collection de M. le comte Dejean. VI. Paris. 804 pp. (PDF)
Babington CC. 1841. Dytiscidae darwinianae; or, descriptions of the species of Dytiscidae collected by Charles Darwin, Esq., M.A., Sec. G.S. &C, in South America and Australia, during his voyage in H.M.S. Beagle. Transactions of the Entomological Society of London 3: 1-17. (PDF)
Balfour-Browne F. 1944. New names and new synonymies in the Dytiscidae (Col.). Annals and Magazine of Natural History (Series 11) 11: 345-359.
Balfour-Browne J. 1943. A new generic name in the Dytiscidae (Coleoptera). Proceedings of the Royal Entomological Society of London (B) 12: 172.
Balfour-Browne J. 1944. New names and new synonymies in the Dytiscidae (Col.). Annals and Magazine of Natural History 78: 346-359.
Balfour-Browne J. 1948. The aquatic Coleoptera of Newfoundland and Nova Scotia. The Canadian Entomologist 80: 156-165.
Bergstrasser JAB. 1778. Nomenklatur und Beschreibung der Insekten in der Grafschaft Hana-Münzenberg, wie auch der Wetterau und der angrænzenden Nachbarschaft dies- und jenseits des Mains. Jahrgang I. 88 pp. + 14 pls.
Blatchley WS. 1910. The Coleoptera or beetles of Indiana. Bulletin of the Indiana Department of Geology and Natural Resources, no. 1. 1385 pp. (PDF)
Boisduval JBA, Lacordaire JT. 1835. Faune entomologique des environs de Paris: ou, Species ge´ne´ral des insectes qui se trouvent dans un rayon de quinze a` vingt lieues aux alentours de Paris. Volume 1. Paris. 696 pp. (PDF)
Brown WJ. 1930a. New species of Coleoptera I. The The Canadian Entomologist 62: 87-92.
Brown, W. J. 1930b. Coleoptera of the north shore of the Gulf of the St. Lawrence. The The Canadian Entomologist 62: 231-237.
Casey TL. 1884. Contributions to the descriptive and systematic Coleoptera of North America. Vol. 8, 198 pp. Philadelphia.
Clairville JP, de. 1806. Entomologie helvétique. Volume 2. Zurich. 247 pp. (PDF)
Clark H. 1862. On the Mexican species of Hydropori. The Annals and Magazine of Natural History (Series 3) 10: 173-184. (PDF).
Clark H. 1864. Notes on the genus Hydaticus (Leach), with descriptions of new species. Transactions of the Entomological Society of London (Series 3) 2: 209-222.
Crotch GR. 1873. Revision of the Dytiscidae of the United States. Transactions of the American Entomological Society 4: 383-424. VIEW
Crotch GR. 1874. Descriptions of new species of Coleoptera from the Pacific Coast of the United States. Transactions of the American Entomological Society 5: 73-80. (PDF)
Curtis J. 1827. British Entomology; being illustrations and descriptions of the genera of insects found in Great Britain and Ireland. Volume 4. London, pis. 147-194.
DeGeer C. 1774. Memoires pour servir a l'histoire des insectes, 7. Stockholm. 950 pp.
Dejean PFMA. 1833. Catalogue des Coléoptères de la collection de M. le compte Dejean. Livre 1 and 2. Paris. 176 pp.
Erichson WF. 1832. Genera dyticeorum. (Dissertatio inauguralis). C. Hold, Berolini. [4] + 48 pp. (PDF).
Erichson WF. 1837. Die käfer der mark Brandenburg. Berlin. 740 pp. (VIEW)
Fabricius JC. 1787. Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus 1. Hafniae [=Copenhagen], 20 + 348 pp. (PDF).
Fall HC. 1917. New Dytiscidae. Journal of the New York Entomological Society 25: 163- 182.
Fall HC. 1919. The North American species of Coelambus. Mount Vernon, New York. 20 pp.
Fall HC. 1922. A review of the North American species of Agabus. Mount Vernon, New York. 36 pp. PDF
Fall HC. 1923. A revision of the North American species of Hydroporus and Agaporus. Privately printed. 129 pp.
Fall HC. 1926. A list of the Coleoptera taken in Alaska and adjacent parts of the Yukon Territory in the summer of 1924. Pan-Pacific Entomologist 2: 127-154.
Fall HC. 1927. New Coleoptera XII. The The Canadian Entomologist 59: 136-141.
Fall HC. 1937. A new Agaporus (Dytiscidae-Coleoptera). Entomological News 48: 10-12.
Fery H, Nilsson AN. 1993. A revision of the Agabus chalconatus- and erichsoni-groups (Celeoptera: Dytiscidae). Entomologica scandinavica 24: 79-108.
Folkerts GW. 1978. A preliminary checklist of the Hydadephaga (Coleoptera) of Alabama. The Coleopterists Bulletin 32: 345-347.
Franciscolo ME. 1979. On a new Dytiscidae from a Mexican cave - a preliminary description (Coleoptera). Fragmenta Entomologica Roma 15(1): 233–241.
Gebler FA, von. 1832. Notice sur les Coléoptères qui se trouvent dans le district des mines de Nertchinsk dans la Sibérie Orientale, avec la description de quelques espèces nouvelles. Nouveaux Mémoires de la Société Impériale des Naturalistes de Moscou 2: 23-78.
Gemminger M, von Harold E. 1868. Catalogus coleopterorum hucusque synonymicus et systematicus, Volume 2. Monachii. pp. 425-752. (VIEW)
Gemminger M, Harold E, von. 1868. Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tomus II. Dytiscidae, Gyrinidae, Hydrophilidae, Staphylinidae, Pselaphidae, Gnostidae, Paussidae, Scydmaenidae, Silphidae, Trichopterygidae, Scaphidiidae. E.H. Gummi, Manachii, pp. 425-752 + [6]. (PDF)
Germar EF. 1924. Insectorum species novae aut minus cognitae, descriptionibus illustrate. Impensis J.C. Hendelii et filii, Halae [=Schwäbisch Hall, Württemberg?], 624 pp.
Gordon RD. 1981. New species of North American Hydroporus, niger-tenebrosus group (Coleoptera: Dytiscidae). The Pan-Pacific Entomologist 57:105-123.
Gordon RD, Post RL. 1965. North Dakota water beetles. North Dakota Insects 5: 1-53.
Gozis M, des. 1914. Tableaux de détermination des dytiscides, notérides, hyphydrides, hygrobiides et haliplides de la faune Franco-Rhénane (part). Miscellanea Entomologica 21: 97-112, 113-128, 161-176, 194-216 (separate pagination).
Gschwendtner L. 1922. Beiträge zur Kenntnis der Dytiscidae Transbaikaliens. Weiner entomologische Zeitung 39: 92-94.
Guignot F. 1931. Sur la systématique des Agabus (Col. Dytiscidae). Bulletin de la Société Entomologique de France 36: 201–203.
Guignot F. 1939. Contribution à l’étude des Bidessus. Bulletin de la Société d’Étude des Sciences Naturelles de Vaucluse 10: 51-61.
Guignot F. 1950. Trente-deuxième note note sur les Hydrocanthares. Revue française d'Entomologie 17: 97-104.
Guignot F. 1952. Description de dytiscides inédits de la collection Régimbart et de quelques autres espèces et variétés nouvelles. Revue Française d’Entomologie 19: 166–171.
Gyllenhal L. 1808. Insecta Suecica descripta, Classis I, Coleoptera sive Eleutherata, Tomus 1, pars 1. F.J. Leverentz, Scaris [=Skara, Sweden]. 660 pp. (PDF).
Gylllenhal L. 1813. Insecta Suecica descripta, Classis I, Coleoptera sive Eleutherata, Tomus 1, pars 3. F.J. Leverentz, Scaris =[Skara, Sweden]. 730 pp. (PDF).
Gyllenhal L. 1827. Insects Suecica descripta, Classis I, Coleoptera sive Eleutherata, Tomus 1, pars 4. Lipsiae [=Leipzig]. 761 pp. (PDF).
Harris TW. 1828a. Contributions to Entomology No. V. New England Farmer 7: 156.
Harris TW. 1828b. Contributions to entomology. No. VI. New England Farmer 7: 164.
Harris TW. 1829. Corrections and additions for the "contributions to entomology". New England Farmer 8: 1-2.
Hatch MH. 1928. Studies on Dytiscidae. Bulletin of the Brooklyn Entomological Society 23: 217-229.
Hatch MH. 1933. Studies on Hydroporus. Bulletin of the Brooklyn Entomological Society 28: 21-27.
Hatch MH. 1953. The beetles of the Pacific Northwest. Part 1: Introduction and Adephaga. University of Washington Publications in Biology 16. 340 pp.
Hilsenhoff WL. 1975. Notes on Nearctic Acilius (Dytiscidae), with the description of a new species. Annals of the Entomological Society of America 68: 271-274.
Hilsenhoff WL. 1980. Coptotomus (Coleoptera: Dytiscidae) in eastern North America with descriptions of two new species. Transactions of the American Entomological Society 105:461-471.
Horn GH. 1883. Miscellaneous notes and short studies on North American Coleoptera. Transactions of the American Entomological Society 10: 269-312.
Hummel A. -D. 1823. Observations sur les insectes de 1823. Monographia pelophilarum. Novae species. Essais entomologiques 3. Imprimerie de la Chancellerie privée du Ministère de l'lntérieur, St. Petersbourg. 48 pp.
Kamiya K. 1938. A systematic study of the Japanese Dytiscidae. Journal Tokyo Nogyo Daigaku 5: 1-68.
Kincaid, T. 1900. The metamorphoses of some Alaska Coleoptera. Proceedings of the Washington Academy of Sciences 2: 367-388.
Kirby W. 1837. The Insects. in Richardson J, Fauna boreali-americana, or, The zoology of the northern parts of British America: containing descriptions of the objects of natural history collected on the late northern land expeditions, under command of Captain Sir John Franklin, R.N., Part 4. London. xxxix + 249 pp. VIEW
Kunze G. 1818. Beitrage zur Monographie der Rohrkafer. Neue Schrift, Naturforschenden Gesellschaft zu Halle 2: 57-70.
Laporte FL, de (Comte de Castelnau). 1840. Histoire naturelle des animaux articulés, annelides, crustacés, arachnides, myriapodes et insectes. Coléoptères. M. Duménil, Paris. cx + 324 pp. (PDF)
Larson DJ. 1975. The predaceous water beetles (Coleoptera: Dytiscidae) of Alberta: systematics, natural history and distribution. Questiones Entomologicae 11: 245-406.
Larson DJ. 1991. Revision of North American Agabus Leach (Coleoptera: Dytiscidae): elongatus-, zetterstedti-, and confinis-groups. The The Canadian Entomologist 123(6): 1239-1317.
Larson DJ, Wolfe RW. 1998. Revision of North American Agabus (Coleoptera: Dytiscidae): the semivittatus-group. The Canadian Entomologist 130: 27-54.
Larson DJ, Roughley RE. 2000. Part 2. Adults, pp. 34-833 in Larson DJ., Alarie Y., Roughley RE, Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa, Ontario, Canada. 982 pp.
Leach WE. 1817. The Zoological Miscellany; being descriptions of new or interesting animals. Volume 3. London. 151 pp.
LeConte JL. 1850. IV. General remarks upon Coleoptera of Lake Superior, pp. 201-242 in Agassiz L, Lake Superior: Its physical character, vegetation and animals, compared with those of other and similar regions. Gould, Kendall and Lincoln: Boston. 428 pp. + 8 pls.
LeConte JL. 1852. Descriptions of new species of Coleoptera from California. Annals of the Lyceum of Natural History of New York 5: 125-216. VIEW.
LeConte JL. 1855. Analytical table of the species of Hydroporus found in the United States, with descriptions of new species. Proceedings of the Academy of Natural Sciences of Philadelphia 7: 290-299.
LeConte JL. 1857. Zoological Report No. 1. Report upon insects collected on the survey. Reports of explorations and surveys to ascertain the most practicable and economical route for a railroad from the Mississippi River to the Pacific Ocean. Made under the direction of the Secretary of War, in 1853-5, according to Acts of Congress of March 3, 1853, May 31, 1854, and August 5, 1854. Volume XII, Book II. Washington. Pp. 1-72, pi. 1 & 2.
LeConte JL. 1859a. The Coleoptera of Kansas and eastern New Mexico. Smithsonian Contributions to Knowledge 11: 1-66.
LeConte JL. 1862. Synopsis of the species of Colymbetes inhabiting America north of Mexico. Proceedings of the Academy of Natural Sciences of Philadelphia 14: 521-523.
LeConte JL. 1863. New species of North American Coleoptera. Smithsonian Miscellaneous Collections, no. 167. 86 pp.
LeConte JL. 1866. Additions to the coleopterous fauna of the United States. No. 1. Proceedings of the Academy of Natural Sciences of Philadelphia 18: 361-394.
LeConte JL. 1878. Descriptions of new species, in Hubbard and Schwarz, The Coleoptera of Michigan. Proceedings of the American Philosophical Society 17: 593-626.
Leech HB. 1939. On some Nearctic species of Agabus, with the description of a new species. (Coleoptera: Dytiscidae). The Canadian Entomologist 71: 217-221.
Leech HB. 1941. The species of Matus, a genus of carnivorous water beetles (Coleoptera, Dytiscidae). The Canadian Entomologist 73: 77-83.
Leech HB. 1944. Laccophilus shermani, a new species of water beetle from Arizona and Texas (Coleoptera: Dytiscidae). Entomological News 55: 4-6.
Leech HB. 1948. Contributions toward a knowledge of the insect fauna of lower California, No. 11. Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Limnebiidae. Proceedings of the California Academy of Sciences 25: 348-420.
Linnaeus C. 1758. Systema naturae per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. ed. 10. Vol. 1. Holmiae. 823 pp.
Mannerheim CG. 1852. Zweiter Nachtrag zur Kaefer-fauna der Nord-Amerikanischen Laender des Russischen Reiches. Bulletin de la Société Imperiale des Naturalistes de Moscou 25: 283- 387. PDF
Mannerheim CG. 1853. Dritter Nachtrag zur Kaefer-fauna der Nord-Amerikanischen Laender des Russischen Reiches. Bulletin de la Société Imperiale des Naturalistes de Moscou 26: 95-273.
Melsheimer FE. 1844. Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 2: 26-47.
Miller KB. 1998. Revision of the Nearctic Liodessus affinis (Say 1823) species group (Coleoptera: Dytscidae, Hydroporinae, Bidessini). Entomologica scandinavica 29(3):281-314.
Motschulsky V. 1852. Nouveautés. Études Entomologiques 1: 77-80.
Motschulsky V. 1853. Hydrocanthares de la Russie. Helsingfors. 3-15 pp.
Motschulsky V. 1859. Coléoptères nouveaux de la Californie. Bulletin de la Société Imperiale des Naturalistes de Moscou 2: 122-185.
Motschulsky, V. 1860. Coléoptères de la Sibérie orientale et en particulier des rives de l'Amour, pp. 77-257, 6 pls., 1 map in von Schrenck LV, Reisen und Forschungen im Amur-Lande. Band II, Lieferung 2 Coleopteren. Kaiserlichen Akademie der Wissenschften, St. Petersburg.
Müller OF. 1776. Zoologiae Danicae prodromus: seu Animalium Daniae et Norvegiae indigenarum; characteres, nomina, et synonyma imprimis popularium. Havniae [=Copenhagen], typis Hallageriis. (PDF).
Nilsson AN, Larson DJ. 1984. A reexamination of some Dytiscidae (Coleoptera) from Newfoundland and adjacent areas. Annales Entomoilogici Fennici 50 (1984): 123-125.
Paykull G. 1798. Fauna Suecica: Insecta, Vol. 1. Upsaliae. 358 pp. (PDF)
Régimbart M. 1895. Dytiscides trouvés dans les Tabacs par les soins de M. Antoine' Grouvelle. Annales de la Société entomologique de France 64: 321-348.
Régimbart M. 1906. Voyage de M. Ch. Alluaud dans l'Afrique orientale. Dytiscidae, Gyrinidae, Hydrophilidae. Annales de la Société entomologique de France 75: 235-278.
Sahlberg CR. 1826. Dissertatio entomologica insecta Fennica enumerans. Tomus I, pars XII. Aboae: Frenckel, pp. 169-184.
Sahlberg JR. 1889. För faunan nya Coleoptera. Meddelanden af Societas pro Fauna et Flora Fennica 15 [1888-1889]: 221-222.
Say T. 1823a. Descriptions of Coleopterous insects collected in the late expedition to the Rocky Mountains performed by order of Mr. Calhoun, Secretary of War, under the command of Major Long. Journal of the Academy of Natural Sciences of Philadelphia 3: 139-216.
Say T. 1823b. Descriptions of insects of the families Carabici and Hydrocanthari of Latreille, inhabiting North America. Transactions of the American Philosophical Society 2: 1-109.
Say T. 1824. Coleoptera, pp. 268-378 in Narrative of an expedition to the source of St. Peter's River, performed by order of Mr. Calhoun, Secretary of War, under the command of Stephen H. Long, Major U.S.T.E. Volume 2. Philadelphia.
Say T. 1825. Descriptions of new species of Coleopterous Insects inhabiting the United States. Journal of the Academy of Natural Sciences of Philadelphia 5: 160-202.
Say T. 1834. Descriptions of new North American insects and observations on some already described. Transactions of the American Philosophical Society 4: 409-470.
Schaller JG. 1783. Neue insekten beschrieben. Schriften der naturforschenden Gesellschaft zu Halle 1: 217-328.
Seidlitz GKM, von. 1887. Bestimmungs-Tabelle der Dytiscidae und Gyrinidae des europäischen Faunengebietes. Verhandlungen des Naturforschenden Vereines in Brünn 25[1886]: 3-136.
Sharp D. 1882. On aquatic carniverous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society 2: 1-1003.
Sherman JD. 1913. Some habits of the Dytiscidae. Journal of the New York Entomological Society 21: 43-54.
Sahlberg CR. 1824. Dissertatio entomologica Insecta Fennica enumerans. Vol. 1, part II: 153-168. Aboae.
Sahlberg J. 1871. Anteckningar till Lapplands Coleopter-fauna. Notiser ur Sällskapets pro Fauna et Flora Fennica Förhandlingar 11: 385-440.
Sahlberg J. 1875. Enumeratio Coleopterorum carnivororum Fenniae. Notiser ur Sällskapets pro Fauna et Flora Fennica Förhandlingar 14: 41-200.
Sahlberg J. 1888. Bidrag till Nordvestra Sibirieninsektfauna, Coleoptera insamlade under expeditionerna till Obi och Jenesej 1876 och 1877. (I. Cicindelidae, Carabidae, Dyticidae, Hydrophilidae, Gyrinidae, Dryopidae, Georyssidae, Limnichidae, Heróceridae, Staphylinidae och Micropeplidae). Handlingar Kongliga Svenska Vetenskaps-Akademiens 17 (1879) 4: 1-115 + 1 pl.
Schaum HR. 1868. Dytiscidae. Tomus I, Hälfte 2, in Erichson WF 1868-1900, Naturgeschichte der Insekten Deutschlands. 1 Abtheilung Coleoptera. Nicolai, Berlin, pp. 1-144.
Stephens JF. 1828. Illustrations of British Entomology. Mandibulata, Volume 2. London. 200 pp.
Strand E. 1935. Miscellanea nomenclatorica zoologica et palaeontologica. Folia Zoologica et Hydrobiologica 8: 176.
Sturm J. 1835. Deutschlands Fauna in Abbildungen nach der Natur mit Beschriebungen. V. Abtheilung. Die Insecten, Neuntes Bänden. Käfer. Nürnberg. xii + 120 pp. + Tafeln 203-216.
Thompson CG. 1856. Översigt af de arter inom familjen Dytisci, som blifvit anträffade på Skandinaviska hälfon. Kongliga Svenska Vetenskaps-Akademien Handlinger 12 (1854): 177-237.
Thunberg CP. 1784. Novae Insectorum species descriptae. Nova Acta Regiae Societatis Scientiarum Upsaliensis 4: 1-28. Thunberg CP. 1794. Dissertatio entomologica novas insectorum species sistens, part 6. Upsaliae. pp. 107-130.
Wallis JB. 1924. Two new species of Coelambus. The Canadian Entomologist 56: 105-108.
Wallis JB. 1926. Some new Coleoptera. The Canadian Entomologist 58: 89-95.
Wallis, J. B. 1933. Some new Dytiscidae (Coleoptera). The Canadian Entomologist 65: 268-278.
Wolfe GW. 1984. A vevision of the vittatipennis species group of Hydroporus Clairville, subgenus Neoporus Guignot (Coleoptera: Dytiscidae). Transactions of the American Entomological Society 110(3): 389-433.
Young FN. 1940. Description of a new dytiscid from Florida (Coleoptera: Dytiscidae). The Florida Entomologist 23: 28.
Young FN. 1953a. A new species of Bidessus from southern Michigan (Coleoptera: Dytiscidae). Bulletin of the Brooklyn Entomological Society 48: 111-112.
Young FN. 1953b. The types of Hydradephaga in the W. S. Blatchley Collection, with generic reassignments and synonymies (Coleoptera: Noteridae, Dytiscidae, Gyrinidae, Haliplidae). The Canadian Entomologist 85: 113-119.
Young FN. 1954. The water beetles of Florida. University of Florida Press, Gainsville. ix + 238 pp.
Young FN. 1979. A key to the Nearctic species of Celina with descriptions of new species (Coleoptera: Dytiscidae). Journal of the Kansas Entomological Society 52: 820-830.
Young FN, Severin HC. 1956. Evidence of intergradation between putative species of Colymbetes in South Dakota (Coleoptera: Dytiscidae). Journal of the Kansas Entomological Society 29: 79-83.
Zaitzev P. 1908. Berichtigungen und Zusätze zu den Haliplidae, Dytiscidae und Gyrinidae in den neuesten Katalogen der Coleopteren. Russkoe Entomologicheskoe Obozrenie 7(1907): 114-124.
Zimmermann A. 1919. Die Schwimmkäfer des Deutschen Entomologischen Museums in Berlin-Dahlem. Archive für Naturgeschichte 83: 68-249.
Zimmermann A. 1920. Coleopterorum catalogus, pars 71, Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. Volume 4. Berlin. 326 pp.
Zimmermann A. 1928. Neuer Beitrag zur Kenntnis der Schwimmkäfer. Wiener Entomologische Zeitung 44:165-187.
Zimmermann A. 1934. Monographie der paläarktischen Dytisciden V. Colymbetinae. (1. Teil: Copelatini, Agabini: Gattung Gaurodytes Thorns.). Koleopterologische Rundschau 20: 138- 214.

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July 10, 2017
• Added taxonomy reference section and made numerous corrections to list; separated Dytsicidae to its own page
November 21, 2012
• Beginning to keep track of changes to the web site pages
• Removed individual pages for families into one long list, families accessed from left menu via anchor redirects within page.

Page created: November 01, 2002; Last update December 22, 2017 (EB)